Desmognathus marmoratus (Moore, 1899)
Leurognathus marmorata Moore, 1899, Proc. Acad. Nat. Sci. Philadelphia, 51: 316. Holotype: ANSP 19610, according to Fowler and Dunn, 1917, Proc. Acad. Nat. Sci. Philadelphia, 69: 21; Malnate, 1971, Proc. Acad. Nat. Sci. Philadelphia, 123: 348. Type locality: "large clear rocky pool on the south flank of Grandfather Mt., N.C. [Avery County, North Carolina, USA], and at an elevation of about 3,500 feet".
Leurognathus marmoratus — Brimley, 1907, J. Elisha Mitchell Sci. Soc., 23: 154.
Leurognathus marmorata marmorata — Pope, 1928, Am. Mus. Novit., 306: 14.
Leurognathus marmorata intermedia Pope, 1928, Am. Mus. Novit., 306: 14. Holotype: AMNH 25557, by original designation. Type locality: "Davis Gap, Waynesville, North Carolina", USA; emended by Bishop, 1943, Handb. Salamanders: 224, to "Davis Farm, about 2 miles east and a little north of Waynesville, (Haywood County) on Highway No. 276." Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30.
Leurognathus intermedia — Pope and Hairston, 1947, Fieldiana, Zool., 31: 160.
Leurognathus marmorata aureata Martof, 1956, Occas. Pap. Mus. Zool. Univ. Michigan, 575: 2. Holotype: UMMZ 111566, by original designation. Type locality: "Jarrard's Creek, 0.2 of a mile below its crossing of U.S. Route 19, about 9 air miles north-northeast of Dahlonega, Lumpkin County, Georgia; 1,550 feet elevation", USA. Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30.
Leurognathus marmorata roborata Martof, 1956, Occas. Pap. Mus. Zool. Univ. Michigan, 575: 4. Holotype: UMMZ 111568, by original designation. Type locality: "Reed Creek, along Burrell Ford Road, 0.5 of a mile from its junction with Glade School Road, about 3.5 miles northwest of Pine Mountain Community, Rabun County, Georgia; 2,350 feet elevation", USA. Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30.
Leurognathus marmorata melania Martof, 1956, Occas. Pap. Mus. Zool. Univ. Michigan, 575: 6. Holotype: UMMZ 111564, by original designation. Type locality: "Otter Creek, tributary of the Nantahala River, 0.5 of a mile west of Tellico Gap, Macon County, North Carolina; 3,600 feet elevation", USA. Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30.
Leurognathus marmoratus marmoratus — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 227.
Leurognathus marmoratus aureatus — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 228.
Leurognathus marmoratus intermedius — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 228.
Leurognathus marmoratus melanius — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 229.
Leurognathus marmoratus roboratus — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 228.
Leurognathus marmoratus — Martof, 1962, Am. Midl. Nat., 67: 30. Rejects status of subspecies.
Desmognathus marmoratus — Bernardo, 1994, Am. Nat., 143: 15. Titus and Larson, 1996, Syst. Biol., 45: 451.
English Names
Moore's Triton (Desmognathus marmorata: Brimley, 1907, J. Elisha Mitchell Sci. Soc., 23: 154).
Shovel-nosed Salamander (Desmognathus marmorata: Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 32; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 174; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 270; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 21; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 16; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 25).
Shovelnose Salamander (Desmognathus marmorata: Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 7; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 32; Collins, 1997, Herpetol. Circ., 25: 6; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 12).
Southern Shovel-nosed Salamander (Leurognathus marmorata intermedia [no longer recognized]: Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 32; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 174).
Pope's Salamander (Leurognathus marmorata intermedia) [no longer recognized]: Bishop, 1943, Handb. Salamanders: 224).
Moore's Salamander (Leurognathus marmorata marmorata) [no longer recognized]: Bishop, 1943, Handb. Salamanders: 220).
Northern Shovel-nosed Salamander (Leurognathus marmorata marmorata) [no longer recognized]: Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 32; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 174).
Golden Shovel-nosed Salamander (Leurognathus marmorata aureatus [no longer recognized]: Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 228).
Husky Shovel-nosed Salamander (Leurognathus marmorata roboratus [no longer recognized]: Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 228).
Black Shovel-nosed Salamander (Leurognathus marmorata melanius [no longer recognized]: Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 229).
Distribution
Southwestern Virginia and southward into the Great Smoky Mountains of western North Carolina and eastern Tennessee to northeastern Georgia and the northwestern tip of South Carolina, USA.
Comment
Reviewed by Martof, 1963, Cat. Am. Amph. Rept., 3: 1-2 (as Leurognathus marmoratus). Voss, Smith, Beachy, and Heckel, 1995, J. Herpetol., 29: 493-497, reported on geographic allozyme variation and suggested that taxonomic revision (likely recognition of additional species) is warranted. Jones, Voss, Ptacek, Weisrock, and Tonkyn, 2006, Mol. Phylogenet. Evol., 38: 280-287, and Wooten and Rissler, 2011, Acta Herpetol., Firenze, 6: 175-208, suggested that neither Desmognathus marmoratus nor Desmognathus quadramaculatus are monophyletic, instead being composed of multiple inter-related species.
External Links
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- For additional sources of information from other sites search Google
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- To search the NIH genetic sequence database, see GenBank
- For information aggregation from other sites and some original accounts see AmphibiaWeb report
- For further information on conservation status and distribution see the IUCN Redlist
- For related information on conservation and images as well as observation see iNaturalist;
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