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Lithobates warszewitschii (Schmidt, 1857)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Lithobates

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Ixalus warszewitschii Schmidt, 1857, Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 24: 11. Holotype: KM 1006/1338, according to Savage, 1970, Proc. California Acad. Sci., Ser. 4, 38: 273-288. Type locality: "Unweit des Vulcanes Chiriqui, zwischen 6000′ und 7000′ Höhe [Polish feet, therefore = 1370-1600 m according to Savage, 1970, Proc. California Acad. Sci., Ser. 4, 38: 273-288], in einem feuchten, nie trockenen Klima von 12-14°R", Panama.

Ixalus warschewitschii — Schmidt, 1858, Denkschr. Akad. Wiss. Wien, Math. Naturwiss. Kl., 14: 241. Incorrect subsequent spelling. Seemingly another description as new of the same taxon noted by this author in 1857.

Rana coeruleopunctata Steindachner, 1864, Verh. Zool. Bot. Ges. Wien, 14: 264. Syntypes: NHMW 20840.1-4, according to Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 31. Type locality: "Fundort unbekkant". Synonymy by Barbour, 1925, Occas. Pap. Boston Soc. Nat. Hist., 5: 155-156; Taylor, 1952, Univ. Kansas Sci. Bull., 35: 896.

Ranula chrysoprasina Cope, 1866, Proc. Acad. Nat. Sci. Philadelphia, 18: 129. Holotype: In USNM according to Cope, but not listed by Cochran, 1961, Bull. U.S. Natl. Mus., 220. Type locality: "Arriba [= roughly the Meseta Central], Costa Rica". Type locality commented on by Savage, 1974, Rev. Biol. Tropical, 22: 77. Synonymy by Taylor, 1952, Univ. Kansas Sci. Bull., 35: 896.

Ranula coeruleopunctata — Cope, 1866, Proc. Acad. Nat. Sci. Philadelphia, 18: 130.

Hylarana coeruleopunctata — Steindachner, 1867, Reise Österreichischen Fregatte Novara, Zool., Amph.: 48.

Trypheropsis chrysoprasinus — Cope, 1868, Proc. Acad. Nat. Sci. Philadelphia, 20: 117.

Rana caeruleopunctata — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 50. Incorrect subsequent spelling.

Rana chrysoprasina — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 49.

Hylarana chrysoprasina — Brocchi, 1882, Miss. Scient. Mex. Amer. Centr., Rech. Zool., 3(2, livr. 2): 65.

Chilixalus Warszewiczii — Werner, 1899, Zool. Anz., 22: 117. Incorrect subsequent spelling of the species name. Considered to be the coining of new species name by Dubois, 1999, Alytes, 17: 85, although Werner (p. 117) makes it clear that he is examining the type of Ixalus warszewitschii Schmidt (although he misspells the species name).

Rana zeteki Barbour, 1925, Occas. Pap. Boston Soc. Nat. Hist., 5: 156. Holotype: MCZ 10031, by original designation. Type locality: "Barro Colorado Island, Gatun Lake, Canal Zone of Panama". Synonymy by Dunn, 1931, Occas. Pap. Boston Soc. Nat. Hist., 5: 416.

Rana warschewitschii zeteki — Dunn, 1931, Occas. Pap. Boston Soc. Nat. Hist., 5: 416.

Rana warschewitschii warschewitschii — Dunn, 1931, Occas. Pap. Boston Soc. Nat. Hist., 5: 416.

Rana warszewitschii — Harding, 1983, Cat. New World Amph.: 55. Hillis and de Sá, 1988, Herpetol. Monogr., 2: 3.

Rana (Rana) warszewitschii — Dubois, 1987 "1986", Alytes, 5: 42. by implication.

Rana (Trypheropsis) warszewitschii — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 330.

Rana (Novirana, Sierrana, Ranula, Levirana, Trypheropsis) warszewitschii — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 305. See Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, and Dubois, 2007, Cladistics, 23: 390-402, for relevant discussion of nomenclature. Invalid name formulation under the International Code of Zoological Nomenclature (1999) as discussed by Dubois, 2007, Cladistics, 23: 395.

Lithobates warszewitschii — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369. Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1-13; by implication.

Lithobates (Lithobates) warszewitschii — Dubois, 2006, C. R. Biol., Paris, 329: 829. Dubois, 2006, Mol. Phylogenet. Evol., 42: 325.

Rana (Trypheropsis) warszewitschii — Hillis, 2007, Mol. Phylogenet. Evol., 42: 335-336. by implication.

English Names

Warszewitsch's Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Distribution

Humid lowlands on the Atlantic drainage from northeastern Honduras to central Panama, 0-1740 m elevation.

Comment

Sole member of the Rana (Trypheropsis) warszewitschii group of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 330. In the Rana palmipes group of Hillis and de Sá, 1988, Herpetol. Monogr., 2: 1-26. See Taylor, 1952, Univ. Kansas Sci. Bull., 35: 896-899, and Hillis and de Sá, 1988, Herpetol. Monogr., 2: 1-26, for synonymy and discussion. See accounts by Villa, 1990, Cat. Am. Amph. Rept., 459: 1-2, Savage, 2002, Amph. Rept. Costa Rica: 404-405, and McCranie and Wilson, 2002, Amph. Honduras: 494-496. McCranie, 2007, Herpetol. Rev., 38: 38, summarized the departmental distribution in Honduras. See statement of geographic range, habitat, and conservation status (as Rana warszewitschii) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 634. See comments by Sunyer, Páiz, Dehling, and Köhler, 2009, Herpetol. Notes, 2: 189-202, regarding Nicaraguan populations.

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