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Hylarana cubitalis (Smith, 1917)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana

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Rana cubitalis Smith, 1917, J. Nat. Hist. Soc. Siam, 2: 277. Holotype: BMNH 1947.2.2.35, according to Ohler, 2007, Alytes, 25: 67. Type locality: "Doi Nga Chang, N. Siam [Thailand]. . . . Collected on the banks of a small stream,. . . at about 500 metres elevation".

Rana (Hylorana) cubitalis — Boulenger, 1920, Rec. Indian Mus., 20: 124.

Rana (Hylarana) cubitalis — Bourret, 1942, Batr. Indochine: 316-317. Dubois, 1987 "1986", Alytes, 5: 42; by implication.

Rana (Sylvirana) cubitalis — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326.

Rana (Sylvirana) nigrotympanica Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 341. Holotype: Designated as specimen figured in fig. 4-1 in Liu and Hu, 1960 "1959", Acta Zool. Sinica, 11: 508-538; given as CIB 571162 by Ohler, 2007, Alytes, 25: 67. Type locality: Not designated beyond "Chine"; given as "Mengyan (21° 37′ N, 101° 22′ E), Xishuanbann Daizu Zizhizhou, Yunnan Sheng, China" by Ohler, 2007, Alytes, 25: 67, who made the synonymy.

Rana (Hylarana) nigrotympanica — Fei, 1999, Atlas Amph. China: 172.

Hylarana cubitalis — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237. by implication; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3; by implication.

Hylarana nigrotympanica — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237. by implication; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 3; by implication.

Sylvirana cubitalis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370. Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34. See comment under Ranidae record.

Sylvirana nigrotympanica — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370. Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 34. Undiscussed recognition. See comment under Ranidae record.

Hylarana (Sylvirana) nigrotympanica — Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1124.

English Names

Siam Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107).

Yellow Stream Frog (Nutphund, 2001, Amph. Thailand: 115).

Darkeared Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 130).

Black-eared Frog (Hylarana nigrotympanica [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108; Fei, 1999, Atlas Amph. China: 172).

Distribution

Karen Hills of Myanmar into Thailand, also in southeastern Thailand on the Cambodia border; Yunnan, Guangxi, and Hainan I., China; northern Laos; presumably into adjacent Vietnam.

Comment

Prior to the revision of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, placed in the Section Hylarana, subsection Hydrophylax, subgenus Sylvirana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 326. See Taylor, 1962, Univ. Kansas Sci. Bull., 43: 424-427, and Bourret, 1942, Batr. Indochine: 316-317, for discussion. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 84-86, to compare this species to other ranids of the Sunda Shelf region. Nutphund, 2001, Amph. Thailand: 116, provided a brief characterization and photograph (which Ohler, 2003, Alytes, 21: 101, did not think identifiable beyond Limnonectes sp.). Chan-ard, 2003, Photograph. Guide Amph. Thailand: 130-131, provided a very brief account, map for Thailand, and photograph. Zhao and Adler, 1993, Herpetol. China: 132, regard the status of Rana nigrotympanica questionable, pending examination of specimens. Yang, 1991, Amph. Fauna of Yunnan: 159-161, provided an account as Rana varians. Fei, 1999, Atlas Amph. China: 172-173, provided a brief account, map, and figure as Hylarana nigrotympanica. Stuart, Heatwole, and Tan, 2006, Hamadryad, 30: 108-113, provided a record, as Rana nigrotympanica, for northern Laos and discussed the nomenclatural confusions with Rana varians. See discussion by Ohler, 2007, Alytes, 25: 55-74. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 67-68, provided a brief account (as Rana cubitalis) for Yunnan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1124-1128, provided an account, figures, and map for China and placed it in their Hylarana (Sylvirana) nigrotympanica group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 311 (as Hylarana cubitalis), 318 (as Hylarana nirotympanica) provided brief accounts. Zhang and Wen, 2000, Amph. Guangxi: 88, provided an account (as Rana varians) for Guangxi. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 74-75, provided a brief account (as Rana varians) for Yunnan, China. Shi, 2011, Amph. Rept. Fauna Hainan: 105-106, provided an account for Hainan as Rana nigrotympanica.

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