Amphibian Species of the World 5.6, an Online Reference

Engystoma ustum Cope, 1866, Proc. Acad. Nat. Sci. Philadelphia, 18: 131. Holotype: USNM 24965 according to Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 188; Cochran, 1961, Bull. U.S. Natl. Mus., 220: 48. Type locality: "Guadalaxara [= Guadalajara, Jalisco], West Mexico"; apparently in error, according to Nelson, 1972, J. Herpetol., 6: 119, who corrected the type locality to "Tecoman, Colima, Mexico".

Systoma ustum — Cope, 1867, J. Acad. Nat. Sci. Philadelphia, Ser. 2, 6: 194.

Engystoma mexicanum Peters, 1870 "1869", Monatsber. Preuss. Akad. Wiss. Berlin, 1869: 881. Holotype: ZMB 6667 according to Nelson, 1972, Herpetol. Rev., 4: 90, and Bauer, Günther, and Robeck, 1996, Mitt. Zool. Mus. Berlin, 72: 263. Type locality: "wareren Gegenden Mexicos (Matamoros u. a. o.)" (= warmer parts of Mexico (Matamoros and other areas); stated as "State of Puebla, Mexico" by Parker, 1934, Monogr. Frogs Fam. Microhylidae: 149; restricted to "[Izucar de] Matamoros, Puebla", Mexico, by Taylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 603, and Smith and Taylor, 1950, Univ. Kansas Sci. Bull., 33: 341; this restriction rejected by Nelson, 1972, Herpetol. Rev., 4: 90. Synonymy by Cope, 1871, Proc. Acad. Nat. Sci. Philadelphia, 23: 216; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 161.

Eupemphix gadovii Boulenger, 1903, Ann. Mag. Nat. Hist., Ser. 7, 12: 552. Syntypes: Not stated; BMNH 1903.9.30.259-261 according to Kellogg, 1932, Bull. U.S. Natl. Mus., 160: 188; reregistered 1947.2.11.54-56 according to Nelson, 1972, Cat. Am. Amph. Rept., 123: 1. Type locality: "San Mateo del Mar", near Tehuantepec, Oaxaca, Mexico. Synonymy by Stejneger, 1910, Proc. Biol. Soc. Washington, 23: 165-168.

Gastrophryne usta — Stejneger, 1910, Proc. Biol. Soc. Washington, 23: 166.

Engystoma usta — Nieden, 1926, Das Tierreich, 49: 64.

Microhyla usta — Parker, 1934, Monogr. Frogs Fam. Microhylidae: 126.

Microhyla usta gadovii — Taylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 603.

Microhyla usta usta — Taylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 603.

Microhyla gadovii — Maldonado-Koerdell, 1953, in Beltrán (ed.), Vida Silvestre y Recursos Nat. Largo Carr. Panamericana: 117.

Gastrophryne usta usta — Carvalho, 1954, Occas. Pap. Mus. Zool. Univ. Michigan, 555: 13.

Gastrophryne usta gadovii — Carvalho, 1954, Occas. Pap. Mus. Zool. Univ. Michigan, 555: 13.

Gastrophryne usta gadowi — Duellman, 1960, Univ. Kansas Publ. Mus. Nat. Hist., 13: 67. Incorrect subsequent spelling.

Gastrophryne usta retifera Lynch, 1965, Trans. Kansas Acad. Sci., 68: 397. Holotype: UIMNH 20048, by original designation. Type locality: "near San Andres Tuxtla, Veracruz, Mexico". Status rejected by Nelson, 1972, J. Herpetol., 6: 122.

Hypopachus ustum — Streicher, Cox, Campbell, Smith, and de Sá, 2012, Mol. Phylogenet. Evol., 64: 645-653. Incorrect gender of species name.

Hypopachus ustus — Frost, 2013, Amph. Spec. World Online, vers. 5.6: . Correction of gender.

English Names

Two-spaded Narrowmouth Toad (Liner, 1994, Herpetol. Circ., 23: 22).

Two-spade Narrowmouth Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 89).

Two-spaded Narrow-mouthed Toad (Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 14).

Distribution

Central Sinaloa, the Balsas Depresion, and central Veracruz to Chiapas (Mexico), Guatemala, and El Salvador on the Pacific versant.

Comment

Reviewed by Nelson, 1972, Cat. Am. Amph. Rept., 123: 1-2. Published records for elevations above 1000 m and those for the Lerma-Chapala drainage system are apparently spurious, as discussed by Nelson, 1972, J. Herpetol., 6: 118-119, who also provided details of distribution. Subspecies suggested, most recently by Lynch, 1965, Trans. Kansas Acad. Sci., 68: 396-400; rejected by Nelson, 1972, J. Herpetol., 6: 122. Valenzuela Galván, Chávez-Juárez, and Alcaraz-Cruz, 2004, Herpetol. Rev., 35: 184, provided the Morelos, Mexico, record. Köhler, Vesely, and Greenbaum, 2005 "2006", Amph. Rept. El Salvador: 62-63, provided an account (for El Salvador) and a color photograph.

External Links

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

• For additional sources of information from other sites search Google
• For images search Arkive, CalPhoto Images and Google Images
• To search the NIH genetic sequence database, see GenBank
• For information aggregation from other sites and some original accounts see AmphibiaWeb report
• For further information on conservation status and distribution see the IUCN Redlist
• For related information on conservation and images as well as observation see iNaturalist;
• for a quick link to their maps see iNaturalist KML