Ischnocnema Reinhardt and Lütken, 1862 "1861", Vidensk. Medd. Dansk Naturhist. Foren., Ser. 2, 3: 239. Type species: Leiuperus verrucosus Reinhardt and Lütken, 1862, by monotypy.
Leiyla Keferstein, 1868, Arch. Naturgesch., 34: 296. Type species: Leiyla guentherii Keferstein, 1868, by monotypy. Also described as new by Keferstein, 1868, Nachr. Ges. Wiss. Göttingen, 1868: 330. Synonymy with Hylodes sensu lato by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 198; with Eleutherodactylus by Savage, 1974, Herpetologica, 30: 289-299; with Ischnocnema by Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.
Liyla — Cope, 1870 "1869", Proc. Am. Philos. Soc., 11: 160. Incorrect subsequent spelling of Leiyla Keferstein.
Basanitia Miranda-Ribeiro, 1923, Rev. Mus. Paulista, Săo Paulo, 13: 851. Type species: Basanitia lactea Miranda-Ribeiro, 1923, by monotypy. Synonymy by Lynch, 1968, Copeia, 1968: 875-876. Provisional synonymy with Ischnocnema by Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2..
Phrynanodus Ahl, 1933, Zool. Anz., 104: 29. Type species: Phrynanodus nanus Ahl, 1933, by monotypy. Synonymy with Eleutherodactylus by Lynch, 1968, Copeia, 1968: 875-876. Provisional synonymy with Ischnocnema by Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.
Big-headed Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 81).
Central and southern Brazil; adjacent northern Argentina; possibly into adjacent Paraguay.
Removed from the synonymy of Eleutherodactylus and redelimited on the basis of molecular evidence by Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, 104: 10092-10097, and Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 1-182, where it had been placed by Caramaschi and Canedo, 2006, Zootaxa, 1116: 43. See comment under Brachycephalidae. Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 1-182, defined species series noted in the comments. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, their study of Genbank sequences, confirmed the sister-taxon relationship with Brachycephalus and provided a tree of exemplar species.
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