Vences, M., F. Glaw, J. Kosuch, I. Das, and M. Veith . 2000. Polyphyly of Tomopterna (Amphibia: Ranidae) based on sequences of the mitochondrial 16S and 12S rRNA genes, and ecological biogeography of Malagasy relict amphibian groups. In: Lourenço, W. R., and S. M. Goodman (Ed.), Diversité et Endéémisme à Madagascar/Diversity and Endemism in Madagascar. Société de Biogéographie [Paris, France], 229-242.