Vences, M., J. Kosuch, S. Lötters, A. Widmer, K.-H. Jungfer, J. Köhler, and M. Veith . 2000. Phylogeny and classification of Poison Frogs (Amphibia: Dendrobatidae), based on mitochondrial 16S and 12S ribosomal RNA gene sequences. Molecular Phylogenetics and Evolution 15: 34-40.