Heyer, W. R., and L. R. Maxson . 1982. Distributions, relationships and zoogeography of lowland frogs. The Leptodactylus complex in South America, with special reference to Amazonia. In: Prance, G. T. (Ed.), Biological Diversification in the Tropics. Proceedings of the Fifth International Symposium of the Association for Tropical Biology, held at Macuto Beach, Caracas, Venezuela, February 8-13, 1979. Columbia University Press [New York], 375-388.