Lötters, S., A. Van der Meijden, L. A. Coloma, R. Boistel, P. Cloetens, R. Ernst, E. Lehr, and M. Veith . 2011. Assessing the molecular phylogeny of a near extinct group of vertebrates: the Neotropical harlequin frogs (Bufonidae; Atelopus). Systematics and Biodiversity 9: 45-57.