Bibliography
Köhler, J., D. Koscinski, J. M. Padial, J. C. Chaparro, P. Handford, S. C. Lougheed, and I. De la Riva . 2010. Systematics of the Andean gladiator frogs of the Hypsiboas pulchellus species group (Anura, Hylidae). Zoologica Scripta. Stockholm 39: 572-590.
