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(Female redundant descriptions removed)
Aposphragisma nocturnum       Get publication

            Male (PBI-OON00015995). Total length 2.15. CEPHALOTHORAX: Carapace red-brown, without any pattern, ovoid in dorsal view, pars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, with rounded posterolateral corners, posterolateral edge without pits, posterior margin not bulging below posterior rim, anterolateral corners without extension or projections, posterolateral surface without spikes, surface of elevated portion of pars cephalica smooth, sides finely reticulate, thorax without depressions, fovea absent, denticles at carapace margin very small; carapace margin with sluice, reaching level of coxa I, surface appears slightly wrinkled, without radiating rows of pits; lateral margin straight, rebordered, with blunt denticles; plumose setae near posterior margin of pars thoracica absent; non-marginal pars cephalica setae light, needle-like, present in u-shaped row, setae arranged in a long u-shaped row reaching posterior edge of pars cephalica; each setae originating from the middle of a pitlike structure with partly swollen edges, especially pronounced at posterior edge of pars cephalica, where hair bases may appear denticle-like; non-marginal pars thoracica setae light, needle-like, very scattered; marginal setae light, needle-like, denticles at carapace margin represent modified hair bases: setae originating from anterior base of denticles, pointing anteriorly. Clypeus sinuous in front view, vertical in lateral view, high, ALE separated from edge of carapace by their radius or more, median projection absent; setae present, light, needle-like, a pair of long, needle-like setae on each half of the clypeus, with the inner ones being longer than the outer ones, pointing inwards. Chilum absent. Eyes six, well developed, pLE and PME tiny; PME asymmetric, roundish to ovoid, ALE largest, all eyes oval; posterior eye row straight from above, procurved from front; ALE separated by their radius to diameter, ALE-PLE separated by less than ALE radius, PME touching for less than half their length, PLE-PME separated by less than PME radius. Sternum longer than wide, red-brown, uniform, fused to carapace, median concavity absent, with radial furrows between coxae I-II, II-III, III-IV, radial furrow opposite coxae III absent, surface finely reticulate, without pits, microsculpture covering entire surface, sickle-shaped structures absent, anterior margin with continuous transverse groove, posterior margin extending posteriorly beyond anterior edges of coxae IV as single extension, anterior corner unmodified, distance between coxae approximately equal, lateral margins unmodified, without posterior hump, radial furrows between coxae I-II, II-III and III-IV only very weakly developed, rather appearing as indentions of sternum margin; sternum microsculptures lobed / blossom-shaped; sternum surface setae originating from middle of microsculptures or from smooth margin surface, marginal setae slightly thicker; posterior edge of sternum protruding as single extension, covered with small blunt denticles; setae abundant, light, needle-like, evenly scattered, originating from surface, without hair tufts. Mouthparts: Chelicerae, endites and labium red-brown. Chelicerae straight, anterior face in lateral view with steplike swelling; toothlike projection on distal end of inner margin of anterior face not developed; promargin without teeth; fangs without tooth-like projections, directed posteriorly, shape normal, without prominent basal process, tip unmodified; setae light, needle-like, evenly scattered, anterior face of chelicerae with few, rather evenly scattered setae; paturon distal region unmodified, promargin with row of flattened setae, inner margin unmodified. Labium triangular, fused to sternum, anterior margin deeply incised, same as sternum in sclerotization; subdistal portion with unmodified setae, ventral surface with 5 needle-like setae: two subdistal setae (one on each side of labium incision) and a transverse line of 3 more proximal setae. Endites distally not excavated, anteromedian tip unmodified, posteromedian part unmodified, same as sternum in sclerotization, anterolateral margin of each endite with two long setae with their tips bent and pointing inwards. ABDOMEN: ovoid, without long posterior extension, rounded posteriorly, inter-scutal membrane rows of small sclerotized platelets absent posteriorly; dorsum soft portions white, without color pattern. Book lung covers large, ovoid, without setae, anterolateral edge unmodified, large, ovoid (sides slightly convex). Posterior spiracles connected by groove. Pedicel tube short, ribbed, scuto-pedicel region unmodified, scutum not extending far dorsal of pedicel, plumose hairs absent, matted setae on anterior ventral abdomen in pedicel area absent, scuto-pedicel region with slightly modified hair bases (appearing as small denticles). Dorsal scutum strongly sclerotized, red-brown, without color pattern, covering full length of abdomen, no soft tissue visible from above, not fused to epigastric scutum, middle surface smooth, sides smooth, anterior half without projecting denticles, surface smooth, but bases of setae appearing pitlike, giving a punctate impression. Epigastric scutum strongly sclerotized, surrounding pedicel, not protruding, small lateral sclerites absent. Postepigastric scutum strongly sclerotized, red-brown, long, semicircular, covering nearly full length of abdominal length, fused to epigastric scutum, anterior margin unmodified, with long posteriorly directed lateral apodemes. Spinneret scutum present, incomplete ring. Dorsum setae present, light, needle-like. Epigastric area setae uniform, light, needle-like. Postepigastric area setae present, light, needle-like. Spinneret scutum with fringe of long setae. Dense patch of setae anterior to spinnerets absent. Interscutal membrane with setae. Colulus present. LEGS: orange-brown, without color pattern; femur IV not thickened, same size as femora I-III, patella plus tibia I shorter than carapace, tibia I unmodified, tibia IV ventral scopula absent, metatarsi I and II meso-apical comb absent , metatarsi III and IV weak ventral scopula absent, femora in dorsal view: femora I-II procurved, femur III more or less straight, femur IV recurved, tibiae tibia III & Tibia IV with specialized hairs on ventral apex, metatarsi metatarsi III & IV with a cluster of dagger like hairs on ventral apex with serrated lateral margins; dorsal apex of metatarsi I-IV with small, shieldlike appendage. Leg spines absent. Tarsi I to IV superior claws tooth not examined in detail. Tarsi I to IV without inferior claw. Trichobothria examined with Stereo microscope; tibia: each with three; metatarsus: each with one. GENITALIA: Epigastric region with sperm pore small, oval, situated between anterior and posterior spiracles, rebordered; furrow without omega-shaped insertions, without setae. Palp normal size, not strongly sclerotized, right and left palps symmetrical, proximal segments orange-brown; embolus dark, prolateral excavation absent; trochanter unmodified; femur normal size, one to two times as long as trochanter, without posteriorly rounded lateral dilation, attaching to patella basally; patella shorter than femur, not enlarged, without prolateral row of ridges, setae unmodified; tibia trichobothria with 3 trichobothria; cymbium pale orange, ovoid in dorsal view, fused with bulb but with clearly defined seam between, not extending beyond distal tip of bulb, plumose setae absent, without stout setae, with distal patch of setae; bulb pale orange, 1 to 1.5 times as long as cymbium, stout, spherical, distal part male palp "gamasomorphic"; bulb anterodorsal without conical extension; Bulbus distally with two appendages, the more dorsal embolus and a more ventral second appendage ('conductor'): embolus dark, lamellar, about half as long as bulb, tip pennant-like and membranous; conductor tapering towards rounded tip, about as long as embolus; embolus-conductor-complex mesially bent in dorsal view, fitting into the continuous transverse groove at anterior margin of sternum. Spermophor easily visible using a compound microscope, spermophor-opening lying on retrolateral apex of embolus.
Aposphragisma nocturnum, male abdomen, lateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male abdomen, prolateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male abdomen, posterior view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male carapace, anterior view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male carapace, dorsal view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male abdomen, ventral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male carapace, lateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male carapace, prolateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male habitus, lateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male habitus, ventral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male sternum, ventral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male habitus, dorsal view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male palp, prolateral view (image by MT, specimen ID: ---15995)


Aposphragisma nocturnum, male palp, prolateral view (image by MT, specimen ID: ---15995)




Aposphragisma nocturnum coordinates
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PBI USI Country State Division Locality Sex Type Status Elevation(m) GPS Coordinate Habitat Collection_Date Collector Inst. Code
PBI_OON 00032179  INDONESIA  Kalimantan Timur  Berau District  About 45 km N of Tanjungredeb, 1 km off road Tanjungredeb - Tanjungselor  1 Adult Male  Paratype  190  2.49250; 117.47944  primary forest  29 Sept 2008  P. Schwendinger  MHNG 
PBI_OON 00015995  INDONESIA  Kalimantan Timur    Kutal, Lempake Tanah Merah  1 Adult Male  Holotype    0.50000; 117.15000  rainforest  24 Oct 1979  P. Lehtinen  MZT 

The American Museum of Natural History, New York, in collaboration with
The California Academy of Sciences, San Francisco   The University of California, Berkeley
The Field Museum of Natural History, Chicago   George Washington University, Washington DC
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This material is based upon work supported by the National Science Foundation under Grant No. DEB 0613754. Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation.
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