The World Spider Catalog, Version 4.5
by Norman I. Platnick

© Copyright 2000, 2001, 2002, 2003, 2004 American Museum of Natural History. All Rights Reserved.
Title Page Introduction Families Counts Bibliography Generic List


FAM. MYSMENIDAE Petrunkevitch, 1928

Last updated June 28, 2003

N.B.: removed from the synonymy of Symphytognathidae by Forster & Platnick, 1977: 2; genera reviewed by Brignoli, 1980j: 728; considered a junior synonym of Anapidae by Wunderlich (1986).

Transferred to other families:
Brucharachne Mello-Leitão, 1925c, type B. ecitophila Mello-Leitão, 1925c: 234, f. 2a-d (f, Argentina) -- removed to the Acari (Krantz & Platnick, 1995).
Cepheia Simon, 1894 -- see Synaphridae.
Synaphris Simon, 1894 -- see Synaphridae.

In synonymy:
Lucarachne Bryant, 1940 = Mysmenopsis Simon, 1897 (Platnick & Shadab, 1978a: 5).

Homonym replaced:
Kilifia Baert & Murphy, 1987 -- see Kilifina.

Gen. Acrobleps Hickman, 1979

A. Hickman, 1979: 68, type A. hygrophilus Hickman, 1979; N.B.: described in the Anapidae, transferred here by Platnick & Forster, 1989: 76.

mf hygrophilus Hickman, 1979 *....................Tasmania
A. h. Hickman, 1979: 69, f. 29-33 (Dmf).

Gen. Anjouanella Baert, 1986

A. Baert, 1986: 265, type A. comorensis Baert, 1986.

mf comorensis Baert, 1986 *....................Comoro Is.
A. c. Baert, 1986: 267, f. 1 (Dmf).

Gen. Brasilionata Wunderlich, 1995

B. Wunderlich, 1995i: 545, type B. arborense Wunderlich, 1995.

m arborense Wunderlich, 1995 *....................Brazil
B. a. Wunderlich, 1995i: 545, f. 9-13 (Dm).

Gen. Calodipoena Gertsch & Davis, 1936

N.B.: removed from the synonymy of Mysmena Simon, 1894 by Brignoli, 1980j: 729, contra Levi, 1956a: 3.

Transferred to other genera:
C. tasmaniae (Hickman, 1979) -- see Mysmena.

mf caribbaea (Gertsch, 1960)....................Jamaica, Trinidad
Mysmena c. Gertsch, 1960a: 21, f. 24, 30-31 (Dmf).
C. c. Brignoli, 1980j: 729 (Tmf from Mysmena).

mf colima (Gertsch, 1960)....................Mexico
Mysmena incredula Levi, 1956a: 5, f. 6, 11-14, 17 (mf, misidentified).
Mysmena c. Gertsch, 1960a: 18, f. 25-26, 32-34 (Dmf).
C. c. Brignoli, 1980j: 729 (Tmf from Mysmena).

m conica (Simon, 1895)....................Algeria
Mysmena c. Simon, 1895g: 149 (Dm).
Mysmena c. Brignoli, 1970b: 1404, f. 3-4, 6 (m).
Mysmena c. Wunderlich, 1980d: 267, f. 44-45 (m).
C. c. Brignoli, 1980j: 729 (Tm from Mysmena).

m dumoga Baert, 1988....................Sulawesi
C. d. Baert, 1988: 20, f. 25-26 (Dm).

mf incredula Gertsch & Davis, 1936 *....................USA, Bahama Is., Cuba
C. i. Gertsch & Davis, 1936: 8, f. 32-33 (Df).
Mysmena i. Levi, 1956a: 5, f. 4-5, 8-10, 16, 18-19 (m, Df).
Mysmena i. Gertsch, 1960a: 15, f. 19-23, 28-29 (mf).
C. i. Brignoli, 1980j: 729 (Tmf from Mysmena).
C. i. Baert, 1984a: 604, f. 15-16 (mf).
C. i. Breene et al., 1993: 99, f. 143A-C (mf).

mf mooatae Baert, 1988....................Sulawesi
C. m. Baert, 1988: 18, f. 22-24 (Dmf).

mf stathamae (Gertsch, 1960)....................Mexico, Panama, Jamaica
Mysmena s. Gertsch, 1960a: 20, f. 27, 35-36, 44 (Dmf).
C. s. Brignoli, 1980j: 729 (Tmf from Mysmena).

f tarautensis Baert, 1988....................Sulawesi
C. t. Baert, 1988: 17, f. 21 (Df).

Gen. Calomyspoena Baert & Maelfait, 1983

C. Baert & Maelfait, 1983: 104, type C. santacruzi Baert & Maelfait, 1983.

mf santacruzi Baert & Maelfait, 1983 *....................Galapagos Is.
C. s. Baert & Maelfait, 1983: 104, f. 1-10 (Dmf).

Gen. Crassignatha Wunderlich, 1995

C. Wunderlich, 1995i: 546, type C. haeneli Wunderlich, 1995.

m haeneli Wunderlich, 1995 *....................Malaysia
C. h. Wunderlich, 1995i: 547, f. 14-20 (Dm).

Gen. Iardinis Simon, 1899

N.B.: transferred (as Iardinus, a lapsus) from the Theridiidae to the Symphytognathidae by Gertsch, 1960a: 8; considered incertae sedis by Levi & Levi, 1962: 22 and Forster & Platnick, 1977: 5; provisionally placed here by Brignoli, 1980j: 731.

Transferred to other genera:
I. albulus Gertsch, 1960 -- see Styposis under Theridiidae.
I. boneti (Forster, 1958) -- see Anapistula under Symphytognathidae.

Nomen dubium:
I. weyersi Simon, 1899a: 87, f. 3 (f, Sumatra *) -- Forster & Platnick, 1977: 5.

m martensi Brignoli, 1978....................Nepal
I. m. Brignoli, 1978l: 249, f. 6-7 (Dm).

m mussardi Brignoli, 1980....................India
I. m. Brignoli, 1980j: 733, f. 1-3 (Dm).

Gen. Isela Griswold, 1985

I. Griswold, 1985b: 208, type I. okuncana Griswold, 1985.

mf okuncana Griswold, 1985 *....................South Africa
I. o. Griswold, 1985b: 210, f. 1-30 (Dmf).

Gen. Itapua Baert, 1984

I. Baert, 1984a: 604, type I. tembei Baert, 1984.

mf tembei Baert, 1984 *....................Paraguay
I. t. Baert, 1984a: 606, f. 1-9 (Dmf).

Gen. Kekenboschiella Baert, 1982

K. Baert, 1982: 303, type K. marijkeae Baert, 1982; N.B.: several undescribed females from New Guinea are illustrated by Baert, 1984b.

m awari Baert, 1984....................New Guinea
K. a. Baert, 1984b: 232, f. 12-15 (Dm).

mf marijkeae Baert, 1982 *....................New Guinea
K. m. Baert, 1982: 303, f. 1-11 (Dmf).

m nubiai Baert, 1984....................New Guinea
K. n. Baert, 1984b: 231, f. 9-11 (Dm).

m vangoethemi Baert, 1982....................New Guinea
K. v. Baert, 1982: 306, f. 12-15 (Dm).

Gen. Kilifina Baert & Murphy, 1992

K. n. name Baert & Murphy, 1992: 104 (replacement name for Kilifia Baert & Murphy, 1987: 194, type Kilifia inquilina Baert & Murphy, 1987, preoccupied).

mf inquilina (Baert & Murphy, 1987) *....................Kenya
Kilifia i. Baert & Murphy, 1987: 194, f. 1-7 (Dmf).

Gen. Leviola Miller, 1970

L. Miller, 1970: 155, type L. termitophila Miller, 1970; transferred from the Theridiidae to the Symphytognathidae by Levi, 1972h: 534, placed here provisionally by Brignoli, 19801: 731.

f termitophila Miller, 1970 *....................Angola
L. t. Miller, 1970: 155, pl. L, f. 1-3, pl. LI, f. 1-6 (Df).

Gen. Maymena Gertsch, 1960

M. Gertsch, 1960a: 30, type Nesticus mayanus Chamberlin & Ivie, 1938; transferred here from the Symphytognathidae by Forster & Platnick, 1977: 2.

Transferred to other genera:
M. bruneti Gertsch, 1960 -- see Plato under Theridiosomatidae.

mf ambita (Barrows, 1940)....................USA
Theridium ambitum Barrows, 1940: 132, f. 5, 5A (Dm).
M. a. Gertsch, 1960a: 38, f. 54-55, 59, 71-72 (Tm from Theridion, Df).
M. a. Griswold et al., 1998: 63, f. 12A (m).

m calcarata (Simon, 1897)....................St. Vincent
Dipoena c. Simon, 1897d: 863 (Dm).
M. c. Gertsch, 1960a: 38, f. 52-53 (Tm from Dipoena).

mf cascada Gertsch, 1971....................Mexico
M. c. Gertsch, 1971: 94, f. 142-143, 145 (Dmf).

mf chica Gertsch, 1960....................Mexico
M. c. Gertsch, 1960a: 35, f. 56-58, 69-70 (Dmf).

mf delicata Gertsch, 1971....................Mexico
M. d. Gertsch, 1971: 94, f. 140-141, 144 (Dmf).

f grisea Gertsch, 1971....................Mexico
M. g. Gertsch, 1971: 95, f. 148 (Df).

mf mayana (Chamberlin & Ivie, 1938) *....................Mexico
Nesticus mayanus Chamberlin & Ivie, 1938: 134, f. 12-13 (Df).
M. m. Gertsch, 1960a: 30, f. 49-51, 60-64 (Tf from Nesticus, Dm).

mf misteca Gertsch, 1960....................Mexico
M. m. Gertsch, 1960a: 36, f. 68 (Df).
M. m. Gertsch, 1971: 95, f. 146-147 (Dm).

mf rica Platnick, 1993....................Costa Rica
M. r. Platnick, in Eberhard, Platnick & Schuh, 1993: 11, f. 10-13 (Dmf).

f roca Baert, 1990....................Peru
M. r. Baert, 1990b: 17, f. 45-46 (Df).

mf sbordonii Brignoli, 1974....................Mexico
M. s. Brignoli, 1974f: 224, f. 8A-D, 9A-F (Dmf).

Gen. Microdipoena Banks, 1895

N.B.: removed from the synonymy of Mysmena Simon, 1894 by Saaristo, 1978: 125.

mf elsae Saaristo, 1978....................Seychelles
M. e. Saaristo, 1978: 124, f. 255-265 (Dmf).
Mysmena e. Roberts, 1978: 932, f. 65-73 (mf).

mf guttata Banks, 1895 *....................USA to Paraguay
M. g. Banks, 1895d: 85 (Dm).
Mysmena g. Bishop & Crosby, 1926: 177, pl. 22, f. 22 (Dm).
Mysmena g. Levi, 1956a: 8, f. 20-30 (m, Df).
Mysmaena g. Gruia, 1977: 162, f. 12-15 (mf).
M. g. Saaristo, 1978: 125 (Tmf from Mysmena).
M. g. Baert, 1984a: 608, f. 10-14 (m).

mf nyungwe Baert, 1989....................Rwanda
M. n. Baert, 1989: 29, f. 1A-B, 2A-B (Dmf).

m vanstallei Baert, 1985....................Cameroon
M. v. Baert, 1985: 51, f. 13 (Dm).

Gen. Mysmena Simon, 1894

N.B.: transferred from the Theridiidae to the Symphytognathidae by Forster, 1959: 270, here by Forster & Platnick, 1977: 2; not a senior synonym of Microdipoena Banks, 1895 (Saaristo, 1978: 125) or of Calodipoena Gertsch & Davis, 1936 or Tamasesia Marples, 1955 (Brignoli, 1980j: 729, contra Levi, 1956a: 3).

Transferred to other genera:
M. caribbaea Gertsch, 1960 -- see Calodipoena.
M. colima Gertsch, 1960 -- see Calodipoena.
M. conica Simon, 1894 -- see Calodipoena.
M. cymbia Levi, 1956 -- see Mysmenopsis.
M. guttata (Banks, 1895) -- see Microdipoena.
M. illectrix Simon, 1895 -- see Mysmenella.
M. incredula (Gertsch & Davis, 1936) -- see Calodipoena.
M. ixlitla Levi, 1956 -- see Mysmenopsis.
M. jobi Kraus, 1967 -- see Mysmenella.
M. quadrimaculata Banks, 1896 -- see Paratheridula under Theridiidae.
M. saltuensis Simon, 1895 -- see Mysmenella.
M. samoensis (Marples, 1955) -- see Mysmenella.
M. stathamae Gertsch, 1960 -- see Calodipoena.

m calypso Gertsch, 1960....................Trinidad
M. c. Gertsch, 1960a: 22, f. 17-18 (Dm).

mf gibbosa Snazell, 1986....................Spain
M. g. Snazell, 1986a: 62, f. 1-8 (Dmf).

m guianensis Levi, 1956....................Guyana
M. g. Levi, 1956a: 7, f. 3 (Dm).

f isolata Forster, 1977....................St. Helena
M. i. Forster, 1977: 129, f. 54a-f (Df).

mf leucoplagiata (Simon, 1879) *....................Southern Europe
Theridion leycoplagiatum Simon, 1879a: 258 (Df).
Dipoena l. Simon, 1881a: 117, pl. 26, f. 2 (f, Dm).
M. l. Simon, 1894a: 558, f. 590 (m).
M. l. Levi, 1956a: 5, f. 1-2 (m).
M. l. Kraus, 1967b: 388, f. 1-11 (mf).
M. l. Loksa, 1973: 283, f. 1-8 (mf).
M. l. Wunderlich, 1980d: 267, f. 24-26 (mf).
M. l. Wunderlich, 1986: 218: f. 227b.

mf phyllicola (Marples, 1955)....................Samoa, Niue
Theridion phyllicolum Marples, 1955a: 488, pl. 57, f. 13, 16, 22, pl. 58, f. 1 (Dmf).
M. p. Forster, 1959: 307 (Tmf from Theridion).

mf tasmaniae Hickman, 1979....................Tasmania
M. t. Hickman, 1979: 74, f. 39-47 (Dmf).
Calodipoena t. Brignoli, 1983c: 376 (Tmf from Mysmena because of alleged relationships with C. incredula Gertsch & Davis, 1936).
M. t. Davies, 1985: 91 (T by Brignoli rejected).

mf vitiensis Forster, 1959....................Fiji
M. v. Forster, 1959: 304, f. 148, 159-166 (Dmf).

f woodwardi Forster, 1959....................New Guinea
M. w. Forster, 1959: 306, f. 147, 167-171 (Df).

Gen. Mysmenella Brignoli, 1980

M. Brignoli, 1980j: 731, type Mysmena illectrix Simon, 1895.

m illectrix (Simon, 1895) *....................Philippines
Mysmena i. Simon, 1895g: 149 (Dm).
M. i. Brignoli, 1980j: 731, f. 5-6 (Tm from Mysmena).

mf jobi (Kraus, 1967)....................Palearctic
Mysmena j. Kraus, 1967b: 392, f. 12-28 (Dmf).
Mysmena j. Shinkai, 1977: 326, f. 12 (f).
M. j. Brignoli, 1980j: 731 (Tmf from Mysmena).
Mysmena j. Wunderlich, 1980d: 267, f. 19-23 (mf).
Mysmena j. Kasal, 1982: 75, f. 6-7 (m).
Mysmena j. Wunderlich, 1986: 222, f. 244.
M. j. Namkung & Lee, 1987: 46, f. 1-8 (mf).
M. j. Thaler & Noflatscher, 1990: 174, f. 31-34 (m).
Mysmena j. Heimer & Nentwig, 1991: 306, f. 822 (mf).
M. j. Namkung, 2002: 146, f. 16.aa-b (mf).

m mihindi Baert, 1989....................Rwanda
M. m. Baert, 1989: 32, f. 1C-D, 2C-D (Dm).

m papuana Baert, 1984....................New Guinea
M. p. Baert, 1984b: 241, f. 28 (Dm).

f saltuensis (Simon, 1895)....................Sri Lanka
Mysmena s. Simon, 1895g: 149 (Df).
M. s. Brignoli, 1980j: 731, f. 4 (Tf from Mysmena).

mf samoensis (Marples, 1955)....................Samoa, Hawaii
Linyphia s. Marples, 1955a: 494, pl. 59, f. 10, 14-16 (Dmf).
Mysmena s. Forster, 1959: 306, f. 148A (Tmf from Linyphia).
M. s. Baert, 1984b: 241, f. 26-27 (Tmf from Mysmena).
M. s. Coddington, 1990: 19, f. 106 (m).

Gen. Mysmeniola Thaler, 1995

M. Thaler, 1995a: 429, type M. spinifera Thaler, 1995.

m spinifera Thaler, 1995 *....................Venezuela
M. s. Thaler, 1995a: 429, f. 1-8 (Dm).

Gen. Mysmenopsis Simon, 1897

N.B.: considered a senior synonym of Lucarachne Bryant, 1940: 349, type L. tibialis Bryant, 1940 by Platnick & Shadab, 1978a: 5; transferred from the Theridiidae to the Symphytognathidae by Gertsch, 1960a: 23, here by Forster & Platnick, 1977: 2.

mf archeri Platnick & Shadab, 1978....................Brazil
M. a. Platnick & Shadab, 1978a: 16, f. 49-53 (Dmf).

mf atahualpa Baert, 1990....................Peru
M. a. Baert, 1990b: 13, f. 27-32 (Dmf).

mf beebei (Gertsch, 1960)....................Trinidad
Lucarachne b. Gertsch, 1960a: 29, f. 47-48 (Dm).
M. b. Platnick & Shadab, 1978a: 9, f. 16-21 (m, Df).

m capac Baert, 1990....................Peru
M. c. Baert, 1990b: 10, f. 13-16 (Dm).

mf cidrelicola (Simon, 1895)....................Venezuela
Theridion c. Simon, 1894a: 536, f. 547, 552, 553 (nomen nudum).
Theridion c. Simon, 1895g: 139 (Dmf).
Lucarachne c. Gertsch, 1960a: 28 (Tm from Theridion; f of Simon belongs to Echinotheridion elicolum).
M. c. Platnick & Shadab, 1978a: 11, f. 28-33 (m, Df).

m cienaga Müller, 1987....................Colombia, Peru
M. c. Müller, 1987e: 185, f. 1-3 (Dm).
M. c. Baert, 1990b: 13, f. 33-39 (m, Df).

mf cymbia (Levi, 1956)....................USA
Mysmena c. Levi, 1956a: 11, f. 38-47 (Dmf).
M. c. Gertsch, 1960a: 25 (Tmf from Mysmena).

mf dipluramigo Platnick & Shadab, 1978....................Panama
Lucarachne beebei Chickering, 1960f: 95, f. 1-4 (f, misidentified).
M. d. Platnick & Shadab, 1978a: 10, f. 22-27 (Dmf).

f femoralis Simon, 1897 *....................St. Vincent
M. f. Simon, 1897d: 865 (Df).
M. f. Gertsch, 1960a: 24, f. 37-39 (f).
M. f. Platnick & Shadab, 1978a: 7, f. 7-9 (f).

mf funebris Simon, 1897....................St. Vincent
M. f. Simon, 1897d: 865 (Df).
M. f. Gertsch, 1960a: 25, f. 40-43 (f, Dm).
M. f. Platnick & Shadab, 1978a: 14, f. 4-6 (m).

mf furtiva Coyle & Meigs, 1989....................Jamaica
M. f. Coyle & Meigs, 1989: 66, f. 4, 24-38 (Dmf).

mf gamboa Platnick & Shadab, 1978....................Panama
M. g. Platnick & Shadab, 1978a: 13, f. 34-39 (Dmf).

mf huascar Baert, 1990....................Peru
M. h. Baert, 1990b: 10, f. 17-26 (Dmf).

mf ischnamigo Platnick & Shadab, 1978....................Panama, Trinidad, Peru
M. i. Platnick & Shadab, 1978a: 7, f. 10-15 (Dmf).
M. i. Baert, 1990b: 17, f. 40-44 (mf).

mf ixlitla (Levi, 1956)....................Mexico
Mysmena i. Levi, 1956a: 9, f. 31, 33-37 (Dmf; not f. 32, see M. mexcala).
M. i. Gertsch, 1960a: 25, f. 45 (Tmf from Mysmena).

mf kochalkai Platnick & Shadab, 1978....................Colombia
M. k. Platnick & Shadab, 1978a: 14, f. 40-45 (Dmf).

m mexcala Gertsch, 1960....................Mexico
Mysmena ixlitla Levi, 1956a: 9, f. 32 (m, misidentified).
M. m. Gertsch, 1960a: 26, f. 46 (Dm).

mf monticola Coyle & Meigs, 1989....................Jamaica
M. m. Coyle & Meigs, 1989: 61, f. 1-3, 8-23 (Dmf).

f pachacutec Baert, 1990....................Peru
M. p. Baert, 1990b: 7, f. 7-8 (Df).

mf palpalis (Kraus, 1955)....................Mexico, Honduras
Lucarachne p. Kraus, 1955b: 30, f. 79-85 (Dmf).
Lucarachne p. Forster, 1959: 328, f. 149.
Lucarachne p. Wunderlich, 1978a: 29, f. 10-12 (mf).
M. p. Platnick & Shadab, 1978a: 18, f. 54-58 (mf).

m penai Platnick & Shadab, 1978....................Ecuador
M. p. Platnick & Shadab, 1978a: 16, f. 46-48 (Dm).

f schlingeri Platnick & Shadab, 1978....................Peru
M. s. Platnick & Shadab, 1978a: 20, f. 68-71 (Df).

mf tengellacompa Platnick, 1993....................Costa Rica
M. t. Platnick, in Eberhard, Platnick & Schuh, 1993: 8, f. 5-9 (Dmf).

mf tibialis (Bryant, 1940)....................Cuba
Lucarachne t. Bryant, 1940: 350, f. 116-120, 123 (Dmf).
M. t. Platnick & Shadab, 1978a: 18, f. 59-64 (mf).
M. t. Coyle & Meigs, 1989: 63, f. 5-7 (f).

mf viracocha Baert, 1990....................Peru
M. v. Baert, 1990b: 5, f. 1-6 (Dmf).

f wygodzinskyi Platnick & Shadab, 1978....................Peru
M. w. Platnick & Shadab, 1978a: 20, f. 65-67 (Df).

mf yupanqui Baert, 1990....................Peru
M. y. Baert, 1990b: 7, f. 9-12 (Dmf).

Gen. Phricotelus Simon, 1895

N.B.: transferred from the Theridiosomatidae to the Symphytognathidae by Levi, 1972b: 534; placed here provisionally by Brignoli, 19801: 731.

f stelliger Simon, 1895 *....................Sri Lanka
P. s. Simon, 1895a: 919, f. 985 (Df).
P. s. Brignoli, 1981a: 14, f. 4 (f).

Gen. Tamasesia Marples, 1955

T. Marples, 1955a: 476, type T. rotunda Marples, 1955; transferred from the Tamasesiidae to the Theridiidae and synonymized with Mysmena Simon, 1894 by Levi, 1956a: 3; removed from synonymy and transferred here by Brignoli, 1980j: 730.

mf acuminata Marples, 1955....................Samoa
T. a. Marples, 1955a: 478, pl. 57, f. 15, 19, 21 (Dmf).

f marplesi Brignoli, 1980....................New Caledonia
T. m. Brignoli, 1980j: 734, f. 7-11 (Df).

mf rotunda Marples, 1955 *....................Samoa
T. r. Marples, 1955a: 476, pl. 57, f. 14, 18, 20, 23 (Dmf).

Gen. Taphiassa Simon, 1880

N.B.: transferred from the Theridiidae to the Symphytognathidae by Levi & Levi, 1962: 29; placed here provisionally by Forster & Platnick, 1977: 2.

Transferred to other genera:
T. punctigera Simon, 1895 -- see Theridiidae incertae sedis.

f impressa Simon, 1880 *....................New Caledonia
T. i. Simon, 1880i: 172 (Df).
T. i. Levi & Levi, 1962: 64, f. 312-313 (f).

Gen. Trogloneta Simon, 1922

N.B.: the spelling Troglonata was a printer's error in the original description; see Bull. Soc. ent. Fr. 1922: 348; transferred from the Theridiidae to the Symphytognathidae by Gertsch, 1960a: 10; placed here by Forster & Platnick, 1977: 2.

mf canariensis Wunderlich, 1987....................Canary Is.
T. canariense Wunderlich, 1987a: 140, f. 381-387 (Dmf).

mf granulum Simon, 1922 *....................Europe
Troglonata g. Simon, 1922: 200 (Dmf).
T. g. Simon, 1926: 313.
T. g. Fage, 1931: 143, f. 11 (mf).
T. g. Levi & Levi, 1962: 64, f. 314-316 (mf).
T. g. Brignoli, 1970b: 1409, f. 1-2, 5 (m).
T. g. Thaler, 1975c: 284, f. 1-22 (mf).
T. g. Wunderlich, 1980d: 267, f. 27-30 (m).
T. g. Wunderlich, 1987a: 140, f. 370-374 (mf).
T. g. Heimer & Nentwig, 1991: 306, f. 821 (mf).
T. g. Breuss, 2001: 187, f. 8-9 (f).

mf madeirensis Wunderlich, 1987....................Madeira
T. madeirense Wunderlich, 1987a: 139, f. 375-380 (Dmf).

mf paradoxa Gertsch, 1960....................USA
T. paradoxum Gertsch, 1960a: 12, f. 11-16 (Dmf).
T. p. Brignoli, 1983c: 380.