The World Spider Catalog, Version 2.0
by Norman I. Platnick

© Copyright 2000, 2001 American Museum of Natural History. All Rights Reserved.
Title Page Introduction Families Counts Bibliography

FAM. CYBAEIDAE Banks, 1892

Last updated Apr. 30, 2001

N.B.: traditionally considered a subfamily of Agelenidae, placed as a subfamily of Dictynidae by Lehtinen, 1967 and elevated to family by Forster, 1970b: 13; considered a senior synonym of Argyronetidae (sub Agelenidae: Cybaeinae) by Grothendieck & Kraus, 1994: 272 (Argyronetidae Thorell 1870 actually has priority over Cybaeidae, but the latter name is now in widespread use).

Transferred to other families:
Amphinecta Simon, 1898 -- see Amphinectidae.
Cambridgea L. Koch, 1872 -- see Stiphidiidae.
Gohia Dalmas, 1917 -- see Desidae.
Urquhartia Bryant, 1933 -- see Matachia under Desidae.

In synonymy:
Bansaia Uyemura, 1938 = Cybaeus L. Koch, 1868 (Yaginuma, 1958f: 76).

Nomen nudum:
Cybaeomyro Kishida, in Komatsu, 1942 (Yaginuma, in Brignoli, 1983c: 484).

Gen. Argyroneta Latreille, 1804

N.B.: transferred here (sub Agelenidae: Cybaeinae) by Grothendieck & Kraus, 1994: 272.

Nomen dubium:
A. palustris Risso, 1826: 165 (France) -- Roewer, 1955c: 1416.

mf aquatica (Clerck, 1757) *....................Palearctic
Araneus aquaticus Clerck, 1757: 143, pl. 6, f. 8 (Dm).
Aranea a. Linnaeus, 1758: 623 (D).
Aranea urinatoria Poda, 1761: 123 (D).
Aranea a. Fabricius, 1775: 436 (D).
Aranea amphibia Müller, 1776: 194 (D).
Aranea a. Fabricius, 1781: 542 (D).
Aranea a. Olivier, 1789: 226 (D).
A. a. Latreille, 1804a: 134 (D).
Aranea a. Latreille, 1804a: 217 (D).
A. a. Sundevall, 1831: 24, 1832: 131 (mf).
A. a. Hahn, 1834: 33, f. 118 (f).
Clubiona fallax Walckenaer, 1837: 603 (Dmf).
A. a. C. L. Koch, 1841a: 60, f. 636 (m).
A. a. Blackwall, 1861a: 137, pl. 8, f. 87 (mf).
A. a. Menge, 1871: 294, pl. 53, f. 170 (mf).
A. a. Simon, 1875a: 29, pl. 5, f. 3 (mf).
A. a. Hansen, 1882: 48, pl. 3, f. 8 (mf).
A. a. Becker, 1896: 184, pl. 15, f. 1 (mf).
A. a. Chyzer & Kulczyn'ski, 1897: 176, pl. 7, f. 11 (mf).
A. a. Simon, 1898a: 234, f. 220-221, 226.
A. a. Bösenberg, 1902: 239, pl. 21, f. 342 (mf).
A. a. Reimoser, 1928: 104, f. 1 (mf).
A. a. Dahl, 1937: 116, f. 34-37 (mf).
A, a. Simon, 1937: 980, 1034, f. 1502-1503 (mf).
A. a. Drensky, 1942: 35, f. 3a-b (mf).
A. a. Crome, 1951: 1, f. 1-21 (mf).
A. a. Locket & Millidge, 1953: 6, f. 2A-B, 4A-B (mf).
A. a. Lehtinen, 1967: 450, f. 268 (m).
A. a. Azheganova, 1968: 20, f. 22-23 (mf).
A. a. Loksa, 1969: 125, f. 84A-B (mf).
A. a. Tyschchenko, 1971: 158, f. 445 (f).
A. a. Miller, 1971: 172, pl. XXVIII, f. 1 (m).
A. a. de Blauwe, 1973a: 4, f. 1-3 (mf).
A. a. Palmgren, 1977a: 8, f. 1.1-4 (mf).
A. a. Paik, 1978e: 302, f. 131.1-2 (mf).
A. a. Zhu, 1982: 29, f. 1-5 (mf).
A. a. Hu, 1984: 212, f. 223.1-4 (mf).
A. a. Roberts, 1985: 154, f. 68a (mf).
A. a. Yaginuma, 1986a: 153, f. 83.1 (mf).
A. a. Song, 1987: 197, f. 155 (mf).
A. a. Chikuni, 1989b: 97, f. 1 (mf).
A. a. Heimer & Nentwig, 1991: 366, f. 951 (mf).
A. a. Bennett, 1992: 6, f. 7-9 (f).
A. a. Grothendieck & Kraus, 1994: 259, f. 1a-b, 9a-c, 10-13, 14b-c, 16 (mf).
A. a. Roberts, 1995: 239, f. (mf).
A. a. Namkung, Kim & Lim, 1996: 112, f. 1-6 (mf).
A. a. Mcheidze, 1997: 202, f. 415-417 (mf).
A. a. Roberts, 1998: 257, f. (mf).
A. a. Song, Zhu & Chen, 1999: 355, f. 207H-J (mf).

Gen. Cedicus Simon, 1875

N.B.: transferred to the Desidae by Lehtinen, 1967: 221; placement uncertain for Forster, 1970b: 21; returned here by Levy, 1996: 118.

f bucculentus Simon, 1889....................Himalayas
C. b. Simon, 1889i: 343 (Df).

f dubius Strand, 1907....................Japan
C. d. Strand, 1907b: 564 (Df).

mf ephthalitus Fet, 1993....................Turkmenistan
C. e. Fet, 1993: 70, f. 1a-e (Dmf).

mf flavipes Simon, 1875 *....................Eastern Mediterranean
C. f. Simon, 1875a: 48, pl. 5, f. 15 (Dmf).
C. f. Simon, 1898a: 245, f. 235-236.
C. f. Kulczyn'ski, 1908c: 77, pl. 2, f. 20-21 (m).
C. f. Lehtinen, 1967: 441, f. 137-139 (mf).
C. f. de Blauwe, 1973a: 8, f. 4-8 (mf).
C. f. Levy, 1996: 118, f. 129-134 (mf).

mf gennadii Fet, 1993....................Turkmenistan
C. g. Fet, 1993: 71, f. 2a-e (Dmf).

mf israeliensis Levy, 1996....................Israel
C. i. Levy, 1996: 120, f. 135-140 (Dmf).

mf maerens Simon, 1889....................Turkmenistan
C. m. Simon, 1889l: 378 (Df).
C. m. Fet, 1993: 73, f. 5a-d (f, Dm).

mf parthus Fet, 1993....................Turkmenistan
C. p. Fet, 1993: 72, f. 4a-e (Dmf).

m pavlovskyi Spassky, 1941....................Tajikistan
C. p. Spassky, 1941: 13, pl. 1, f. 1-2 (Dm).

f pumilus Thorell, 1895....................Myanmar
C. p. Thorell, 1895: 52 (Df).

mf simoni Charitonov, 1946....................Uzbekistan
C. s. Charitonov, 1946: 21, f. 8-9 (Dmf).
C. s. Fet, 1993: 72, f. 3a-c (m).

Gen. Cybaeina Chamberlin & Ivie, 1932

N.B.: placed here by Lehtinen, 1967: 226.

mf confusa Chamberlin & Ivie, 1942....................USA
C. c. Chamberlin & Ivie, 1942a: 19, pl. 3, f. 38 (Df).
C. c. Roth, 1952c: 195, f. 1, 3, 6 (f, Dm).
C. c. Roth & Brame, 1972: 16, f. 21-22 (mf).
C. c. Roth, 1982: 7-8, f. 21-22 (mf).
C. c. Roth, 1985: B1-7, f. 21-22 (mf).
C. c. Roth, 1994: 53, f. 21-22 (mf).

mf minuta (Banks, 1906) *....................USA
Cybaeus minutus Banks, 1906b: 95 (Df).
C. m. Chamberlin & Ivie, 1932: 28, pl. 2, f. 5-8, pl. 3, f. 9-14, pl. 4, f. 28-30 (f, Dm).

mf sequoia Roth, 1952....................USA
C. s. Roth, 1952c: 197, f. 1, 4-5 (Dmf).
C. s. Roth, 1952c: 197, f. 2, 4-5 (Dmf).

f xantha Chamberlin & Ivie, 1937....................USA
C. x. Chamberlin & Ivie, 1937: 225, pl. 9, f. 65 (Df).

Gen. Cybaeota Chamberlin & Ivie, 1933

N.B.: placed here by Lehtinen, 1967: 226.

In synonymy:
C. concolor Chamberlin & Ivie, 1937 = C. nana Chamberlin & Ivie, 1937 (Bennett, 1988: 108).
C. vancouverana Chamberlin & Ivie, 1937 = C. shastae Chamberlin & Ivie, 1937 (Bennett, 1988: 115).
C. wasatchensis Chamberlin & Ivie, 1937 = C. shastae Chamberlin & Ivie, 1937 (Bennett, 1988: 115).

mf calcarata (Emerton, 1911) *....................USA, Canada
Liocranum calcaratum Emerton, 1911: 402, pl. 5, f. 4 (Dmf).
C. c. Chamberlin & Ivie, 1933b: 1, f. 1-10 (mf).
C. calcaratum Kaston, 1977: 37, f. 31-32 (mf).
C. c. Bennett, 1988: 107, f. 19-20, 25-26, 30 (mf).

f munda Chamberlin & Ivie, 1937....................USA
C. m. Chamberlin & Ivie, 1937: 228, pl. 11, f. 83-84 (Df).
C. m. Bennett, 1988: 114, f. 1-2, 33, 35-36 (f).

mf nana Chamberlin & Ivie, 1937....................USA, Canada
C. n. Chamberlin & Ivie, 1937: 229, pl. 10, f. 74-75, pl. 11, f. 79-80 (Dmf).
C. concolor Chamberlin & Ivie, 1937: 227, pl. 11, f. 77-78 (Df).
C. n. Roth & Brame, 1972: 17, f. 24 (m).
C. n. Roth, 1982: 7-8, f. 24 (m).
C. n. Roth, 1985: B1-7, f. 24 (m).
C. n. Bennett, 1988: 108, f. 3-8, 18, 23-24, 34, 37-38 (mf, S).
C. n. Roth, 1994: 53, f. 24 (m).

mf shastae Chamberlin & Ivie, 1937....................USA
C. s. Chamberlin & Ivie, 1937: 227, pl. 10, f. 68-70 (Dmf).
C. wasatchensis Chamberlin & Ivie, 1937: 227, pl. 10, f. 71-73, pl. 11, f. 76 (Dmf).
C. vancouverana Chamberlin & Ivie, 1937: 228, pl. 11, f. 81-82 (Df).
C. s. Roth & Brame, 1972: 17, f. 23 (f).
C. s. Roth, 1982: 7-8, f. 23 (f).
C. s. Roth, 1985: B1-7, f. 23 (f).
C. s. Bennett, 1988: 115, f. 9-17, 21-22, 27-29, 31-32 (mf, S).
C. s. Bennett, 1992: 4, f. 10-11 (f).
C. s. Roth, 1994: 53, f. 23 (f).

Gen. Cybaeozyga Chamberlin & Ivie, 1937

N.B.: placed here by Lehtinen, 1967: 226.

mf heterops Chamberlin & Ivie, 1937 *....................USA
C. h. Chamberlin & Ivie, 1937: 226, pl. 9, f. 66-67 (Dm).
C. h. Roth & Brame, 1972: 18, f. 19, 25-26 (m, Df).
C. h. Roth, 1982: 7-8, f. 19, 25-26 (mf).
C. h. Roth, 1985: B1-7, f. 19, 25-26 (mf).
C. h. Roth, 1994: 53, f. 25-26 (mf).

Gen. Cybaeus L. Koch, 1868

N.B.: considered a senior synonym of Bansaia Uyemura, 1938c: 136, type B. nipponica Uyemura, 1938 by Yaginuma, 1958f: 76 (through transfer of type species; cf. Lehtinen, 1967: 218).

Transferred to other genera:
C. invalidus Simon, 1898 -- see Chresiona under Amaurobiidae.
C. ishikawai (Kishida, 1961) -- see Dolichocybaeus.
C. kirigaminensis Komatsu, 1963 -- see Dolichocybaeus.
C. kuramotoi Yaginuma, 1963 -- see Dolichocybaeus.
C. maculatus Keyserling, 1878 -- see Rhoicinaria under Amaurobiidae.
C. okafujii Yaginuma, 1963 -- see Dolichocybaeus.
C. pirini (Drensky, 1921) -- see Cryphoeca under Hahniidae.
C. varius Keyserling, 1879 -- see Naevius under Amaurobiidae.
C. whanseunensis Paik & Namkung, 1967 -- see Dolichocybaeus.

In synonymy:
C. adenoides Schenkel, 1950 = C. adenes Chamberlin & Ivie, 1932 (Roth, 1956b: 178).
C. grizzlyi Schenkel, 1950 = C. adenes Chamberlin & Ivie, 1932 (Roth, 1956b: 178).
C. patritus Bishop & Crosby, 1926 = C. giganteus Banks, 1892 (Roth & Brown, 1986: 3).
C. silicis Barrows, 1919 = C. giganteus Banks, 1892 (Roth & Brown, 1986: 3).

Nomen dubium:
C. austinensis (Chamberlin, 1924a: 2, f, USA, originally in Parauximus) -- Roth & Brown, 1986: 15.

Nomina nuda:
C. caucasicus Charitonov, 1947 -- see C. abchasicus.
C. limbata Kishida -- Yaginuma, in Brignoli, 1983c: 484.

f abchasicus Charitonov, 1947....................Turkey, Georgia
C. a. Charitonov, 1947b: 18, f. 2 (Dj).
C. caucasicus Charitonov, 1947b: 26 (lapsus in Summary).
C. a. Brignoli, 1978g: 508, f. 79 (Df).

mf adenes Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 24, pl. 6, f. 59 (Df).
C. adenoides Schenkel, 1950b: 88, f. 33 (Df).
C. grizzlyi Schenkel, 1950b: 86, f. 32a-c (Dm).
C. a. Roth, 1956b: 178 (Sm).
C. grizzlyi Bennett, 1992: 4, f. 1-2 (f).

f amicus Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 24, pl. 6, f. 58 (Df).

f angelus Chamberlin & Ivie, 1942....................USA
C. a. Chamberlin & Ivie, 1942a: 16, pl. 5, f. 50 (Df).

mf angustiarum L. Koch, 1868....................Europe to Azerbaijan
C. a. L. Koch, 1868: 47, f. 22-23 (Dmf).
C. tetricus Simon, 1875a: 17, pl. 5, f. 1 (mf, misidentified).
C. tetricus Herman, 1879: 121, 352, pl. 5, f. 107 (mf, misidentified).
C. a. Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 7 (mf).
C. a. Bösenberg, 1902: 231, pl. 21, f. 328 (mf).
C. a. Dahl, 1931: 11, f. 16-17 (mf).
C. a. Simon, 1937: 982, 1034, f. 1506-1507 (mf).
C. a. Drensky, 1942: 36, f. 3c-d (mf).
C. a. Loksa, 1969: 116, f. 79A, E-F (mf).
C. a. Miller, 1971: 173, pl. XXVIII, f. 2-3 (mf).
C. a. de Blauwe, 1973a: 12, f. 9-11 (mf).
C. a. Brignoli, 1977f: 30, f. 6, 9-10 (f).
C. a. Brignoli, 1978e: 282, f. 11 (f, S).
C. angustiorum Heimer & Nentwig, 1991: 360, f. 934 (mf).
C. a. Maurer, 1992: 156, f. 19-22 (mf).

mf aokii Yaginuma, 1972....................Japan
C. a. Yaginuma, 1972c: 25, f. 7-8 (Dmf).
C. a. Yaginuma, 1986a: 143, f. 78.9 (mf).

m aquilonalis Yaginuma, 1958....................China, Japan
C. a. Yaginuma, 1958f: 76, f. 27 (Dm).
C. a. Yaginuma, 1960: 78, f. 71.4 (m).
C. a. Yaginuma, 1971: 78, f. 71.4 (m).
C. a. Yaginuma, 1986a: 143, f. 78.11 (m).

mf aspenicolens Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 21, pl. 6, f. 52-54 (Dmf).

mf balkanus Deltshev, 1997....................Balkans
C. b. Deltshev, 1997b: 1, f. 1-9 (Dmf).

f bam Marusik & Logunov, 1991....................Kurile Is.
C. b. Marusik & Logunov, 1991: 88, f. 1.5, 3.3 (Df).

mf basarukini Marusik & Logunov, 1991....................Sakhalin
C. b. Marusik & Logunov, 1991: 88, f. 1.4, 2.1-2, 3.1-2 (Dmf).

f blasbes Chamberlin & Ivie, 1932....................USA
C. b. Chamberlin & Ivie, 1932: 22, pl. 6, f. 55 (Df).

mf brignolii Maurer, 1992....................Turkey
C. b. Maurer, 1992: 156, f. 27-28 (Dmf).

j broni Caporiacco, 1934....................Karakorum
C. b. Caporiacco, 1934a: 224 (Dj).

f bulbosus Exline, 1935....................USA
C. b. Exline, 1935b: 285, unnumbered f. (Df).
C. b. Exline, 1938: 13, pl. 2, f. 10 (f).

mf cascadius Roth, 1952....................USA
C. c. Roth, 1952b: 212, pl. 1, f. 4-8 (Dmf).
C. c. Roth & Brame, 1972: 20, f. 28 (m).
C. c. Roth, 1982: 7-8, f. 28 (m).
C. c. Roth, 1985: B1-7, f. 28 (m).
C. c. Roth, 1994: 53, f. 28 (m).

f chaudius Exline, 1935....................USA
C. c. Exline, 1935a: 131, f. 3 (Df).
C. c. Exline, 1938: 12, pl. 2, f. 9 (f).

m communis Yaginuma, 1972....................Japan
C. c. Yaginuma, 1972e: 311, f. 37 (Dm).
C. c. Yaginuma, 1986a: 143, f. 78.10 (m).

mf confrantis Oliger, 1994....................Russia
C. c. Oliger, 1994: 51, f. 1a-d, 2a-e (Dmf).

f conservans Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 21, pl. 5, f. 51 (Df; not f. 50).

f consocius Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 25, pl. 6, f. 61 (Df).

f constrictus Chamberlin & Ivie, 1942....................USA
C. c. Chamberlin & Ivie, 1942a: 15, pl. 5, f. 49 (Df).

mf cribelloides Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 26, pl. 6, f. 62 (Df).
C. consocius Gertsch & Ivie, 1936: 22, f. 40 (m, misidentified per Roth, 1956b: 178).
C. c. Chamberlin & Ivie, 1942a: 18, pl. 4, f. 45-46 (Dm).

f cylisteus Zhu & Wang, 1992....................China
C. c. Zhu & Wang, 1992: 342, f. 4-5 (Df).
C. c. Song, Zhu & Chen, 1999: 355,f. 208A-B (f).

mf deletroneus Zhu & Wang, 1992....................China
C. d. Zhu & Wang, 1992: 343, f. 8-12 (Dmf).
C. d. Song, Zhu & Chen, 1999: 355, f. 208C-D, K-L (mf).

f desmaeus Zhu & Wang, 1992....................China
C. d. Zhu & Wang, 1992: 343, f. 6-7 (Df).
C. d. Song, Zhu & Chen, 1999: 355, f. 208E-F (f).

f devius Chamberlin & Ivie, 1942....................USA
C. d. Chamberlin & Ivie, 1942a: 18, pl. 5, f. 48 (Df).

m echinaceus Zhu & Wang, 1992....................China
C. e. Zhu & Wang, 1992: 342, f. 1-3 (Dm).
C. e. Song, Zhu & Chen, 1999: 355, f. 208M-N (m).

mf eutypus Chamberlin & Ivie, 1932....................USA
C. e. Chamberlin & Ivie, 1932: 19, pl. 5, f. 43-45 (Dmf).
C. e. Roth, 1952b: 212, pl. 1, f. 15, pl. 2, f. 17, 33 (mf).

mf exlineae Chamberlin & Ivie, 1937....................USA
C. e. Chamberlin & Ivie, 1937: 224, pl. 9, f. 62-64 (Dmf).
C. e. Roth, 1952b: 216, pl. 2, f. 25-27, 30 (mf).

f fujisanus Yaginuma, 1972....................Japan
C. f. Yaginuma, 1972e: 309, f. 39 (Df).
C. f. Yaginuma, 1986a: 143, f. 78.13 (f).

mf giganteus Banks, 1892....................USA
C. g. Banks, 1892a: 23, pl. 1, f. 71, pl. 5, f. 71 (Df).
C. silicis Barrows, 1919b: 356, pl. 15, f. 7 (Dmf).
C. patritus Bishop & Crosby, 1926: 202, pl. 25, f. 54-55 (Dmf).
C. g. Chamberlin & Ivie, 1932: 14, pl. 4, f. 33 (f).
C. silicis Chamberlin & Ivie, 1932: 15, pl. 4, f. 26-27, 36 (mf).
C. patritus Chamberlin & Ivie, 1932: 15, pl. 4, f. 34-35 (mf).
C. g. Roth & Brown, 1986: 3 (S).

mf gonokawa Ihara, 1993....................Japan
C. g. Ihara, 1993: 118, f. 8-14 (Dmf).

f hatchi Exline, 1935....................USA
C. h. Exline, 1935a: 130, f. 2 (Df).

mf hesper Chamberlin & Ivie, 1932....................USA
C. minor Banks, 1904b: 341, pl. 38, f. 4, pl. 40, f. 44 (Dmf; preoccupied by Chyzer, 1897).
C. minor Chamberlin & Ivie, 1932: 27, pl. 6, f. 63-64 (f).
C. h. Chamberlin & Ivie, 1932: 27, pl. 6, f. 65-66 (Df).
C. minoratus Gertsch, 1936a: 14 (replacement name).

mf hibaensis Ihara, 1994....................Japan
C. h. Ihara, 1994: 87, f. 1-9 (Dmf).

mf hiroshimaensis Ihara, 1993....................Japan
C. h. Ihara, 1993: 116, f. 1-7 (Dmf).

f hystrix Chamberlin & Ivie, 1942....................USA
C. h. Chamberlin & Ivie, 1942a: 18, pl. 5, f. 47 (Df).

mf intermedius Maurer, 1992....................Switzerland, Italy
C. i. Maurer, 1992: 152, f. 5-8, 14, 31 (Dmf).

mf itsukiensis Irie, 1998....................Japan
C. i. Irie, 1998: 97, f. 1-11 (Dmf).

f jaanaensis Komatsu, 1968....................Japan
C. j. Komatsu, 1968b: 24, f. 8, 24, 40, 56, 70, 79, 84 (Df).
C. j. Yaginuma, 1986a: 143, f. 77.7 (f).

f janus Chamberlin & Ivie, 1942....................USA
C. j. Chamberlin & Ivie, 1942a: 15, f. 44 (Df).

mf jilinensis Song, Kim & Zhu, 1993....................China
C. j. Song, Kim & Zhu, 1993: 19, f. 1-4 (Dmf).
C. j. Song, Zhu & Chen, 1999: 355, f. 208G-H, O-P (mf).

mf kiuchii Komatsu, 1965....................Japan
C. k. Komatsu, 1965: 21, f. 1-5 (Dm).
C. k. Komatsu, 1968b: 22, f. 6, 22, 38, 54, 82 (Df).

mf kunashirensis Marusik & Logunov, 1991....................Kurile Is., Japan
C. k. Marusik & Logunov, 1991: 90, f. 2.3-5, 3.4-6 (Dmf).
C. k. Ono & Nishikawa, 1992: 138, f. 1-8 (mf).

f longus Paik, 1966....................Korea
C. l. Paik, 1966b: 33, f. 2, 7-9 (Df).
C. l. Paik, 1978e: 305, f. 132.1-2 (f).

f maculosus Yaginuma, 1972....................Japan
C. m. Yaginuma, 1972e: 312, f. 38 (Df).
C. m. Yaginuma, 1986a: 144, f. 78.12 (f).

f magnus Yaginuma, 1958....................Japan
C. m. Yaginuma, 1958f: 75, f. 28-31 (Df).
C. m. Yaginuma, 1960: 78, f. 71.2 (f).
C. m. Yaginuma, 1971: 78, f. 71.2 (f).
C. m. Yaginuma, 1986a: 143, f. 78.14 (f).

f marinensis Chamberlin & Ivie, 1932....................USA
C. m. Chamberlin & Ivie, 1932: 25, pl. 6, f. 60 (Df).

mf melloteei (Simon, 1886)....................Japan
Cicurina mellotei Simon, 1886d: 58 (Df; patronym for Mellotée).
C. m. Bösenberg & Strand, 1906: 295, pl. 16, f. 457 (f, Dm).
C. m. Yaginuma, 1941: 123, pl. 3.2, f. 2, 2.1-3 (mf).
C. m. Yaginuma, 1960: 78, f. 71.3, 7-8 (mf).
C. m. Yaginuma, 1971: 78, f. 71.3, 7-8 (mf).

mf minor Chyzer, 1897....................Europe
C. m. Chyzer, in Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 9 (Dmf).
C. m. Loksa, 1969: 116, f. 79C, G (mf).
C. m. Brignoli, 1978g: 508, f. 76-78 (mf).
C. m. Polenec, 1985: 103, f. 5-7 (mf).
C. m. Maurer, 1992: 151, f. 9-12, 15, 17 (mf).

f miyosii Yaginuma, 1941....................Japan
C. m. Yaginuma, 1941: 128, f. 1-5 (Df).
C. m. Yaginuma, 1960: 78, f. 71.1 (f).
C. m. Yaginuma, 1971: 78, f. 71.1 (f).
C. m. Yaginuma, 1986a: 143, f. 78.6 (f).

mf montanus Maurer, 1992....................Switzerland, Italy
C. m. Maurer, 1992: 151, f. 1-4, 13, 16, 32 (Dmf).

mf morosus Simon, 1886....................USA, Alaska
C. m. Simon, 1886d: 57 (Df).
C. m. Banks, 1913: 180, pl. 13, f. 37 (f).
C. signifer Emerton, 1923: 240, f. 5d-f (m, misidentified).
C. m. Chamberlin & Ivie, 1932: 20, pl. 1, f. 1-4, pl. 3, f. 15-21 (f, Dm).
C. m. Roth, 1952b: 210, pl. 2, f. 19, 31 (mf).
C. m. Bennett, 1992: 4, f. 3 (f).

mf mosanensis Paik & Namkung, 1967....................Korea
C. nipponicus Paik, 1966b: 31, f. 1, 4-6 (f, misidentified).
C. m. Paik & Namkung, 1967: 22, f. 1-5 (Dmf).
C. m. Paik, 1978e: 306, f. 133.1-7 (mf).

mf multnoma Chamberlin & Ivie, 1942....................USA
C. conservans Chamberlin & Ivie, 1932: 21, pl. 5, f. 50 (not 51) (f, misidentified).
C. m. Chamberlin & Ivie, 1942a: 14, pl. 4, f. 43 (Df).
C. m. Roth, 1952b: 208, pl. 1, f. 1-3, pl. 2, f. 20 (f).
C. m. Roth & Brame, 1972: 20, f. 27 (f).
C. m. Roth, 1982: 7-8, f. 27 (f).
C. m. Roth, 1985: B1-7, f. 27 (f).
C. m. Roth, 1994: 53, f. 27 (f).

mf nipponicus (Uyemura, 1938)....................Japan
Bansaia nipponica Uyemura, 1938c: 137, pls. 12, 13a-g (Dmf).
Bansaia nipponica Yaginuma, 1941: 123, pl. 3.1 (m).
C. n. Yaginuma, 1958f: 76.
C. n. Yaginuma, 1960: 78, f. 71.5-6 (mf).
C. n. Komatsu, 1968b: 22, f. 7, 23, 39, 55, 83 (f).
C. n. Yaginuma, 1971: 78, f. 71.5-6 (mf).
C. n. Yaginuma, 1986a: 143, f. 78.2 (m).
C. n. Chikuni, 1989b: 104, f. 26 (mf).

mf nojimai Ihara, 1993....................Japan
C. n. Ihara, 1993: 123, f. 19-23 (Dmf).

f obedientiarius Komatsu, 1963....................Japan
C. o. Komatsu, 1963a: 9, pl. V, f. A-H (Df).
C. o. Yaginuma, 1986a: 144, f. 77.2 (f).

mf okayamaensis Ihara, 1993....................Japan
C. o. Ihara, 1993: 125, f. 23-26 (Dmf).

f olympiae Exline, 1935....................USA
C. o. Exline, 1935a: 129, f. 1 (Df).
C. o. Exline, 1938: 11, pl. 2, f. 8 (f).

f perditus Chamberlin & Ivie, 1932....................USA
C. p. Chamberlin & Ivie, 1932: 12, pl. 4, f. 31 (Df).

m petegarinus Yaginuma, 1972....................Japan
C. p. Yaginuma, 1972c: 26, f. 9 (Dm).
C. p. Yaginuma, 1986a: 144, f. 78.8 (m).

f potanini Schenkel, 1963....................China
C. p. Schenkel, 1963: 275, f. 156 (Df).
C. p. Song, Zhu & Chen, 1999: 355, f. 208I-J (f).

mf rarispinosus Yaginuma, 1970....................Japan
C. r. Yaginuma, in Yaginuma & Nishikawa, 1970: 137, f. 1-2 (Dmf).
C. r. Yaginuma, Yamaguchi & Nishikawa, 1976: 828, f. 7-9 (mf).

mf raymondi (Simon, 1916)....................France
Cicurina r. Simon, 1916b: 211 (Df).
Cicurina r. Simon, 1937: 1019, 1043, f. 1585 (f).
Cicurina r. de Blauwe, 1973a: 23, f. 22 (f).
C. angustiarum Brignoli, 1978e: 282 (S, rejected).
C. r. Maurer, 1992: 157, f. 25-26, 29-30 (removed mf from S of C. angustiarum).

f reducens Chamberlin & Ivie, 1932....................USA
C. r. Chamberlin & Ivie, 1932: 23, pl. 6, f. 57 (Df).

mf reticulatus Simon, 1886....................USA, Canada, Alaska
C. r. Simon, 1886d: 56 (Dmf).
C. r. Simon, 1898a: 237, f. 234 (m).
C. r. Emerton, 1923: 240, f. 4 (mf).
C. r. Chamberlin & Ivie, 1932: 17, pl. 5, f. 40-42 (mf).
C. r. Exline, 1938: 12, pl. 1, f. 1 (mf).
C. r. Roth, 1952b: 207, pl. 2, f. 18, 22 (mf).

f ryunoiwayaensis Komatsu, 1968....................Japan
C. r. Komatsu, 1968b: 21, f. 9, 25, 41, 57, 77, 85 (Df).

f sanctus (Komatsu, 1942)....................Japan
Cybaeomyro s. Komatsu, 1942b: 65, f. 8 (Df in generic nomen nudum).
Cybaeomyro s. Komatsu, 1961a: 30, f. 19A-C (f; name attributed to Kishida).
C. s. Brignoli, 1983c: 485.
C. s. Yaginuma, 1986a: 146, f. 77.4 (f).

mf scopulatus Chamberlin & Ivie, 1942....................USA
C. s. Chamberlin & Ivie, 1942a: 13, f. 39-40 (Dm).
C. s. Roth, 1952b: 214, pl. 1, f. 13-14, pl. 2, f. 21, 24 (f).

mf septatus Chamberlin & Ivie, 1942....................USA
C. s. Chamberlin & Ivie, 1942a: 17, pl. 4, f. 41-42 (Dmf).
C. s. Roth, 1952b: 219, pl. 2, f. 29 (f).

f shingenni Komatsu, 1968....................Japan
C. s. Komatsu, 1968b: 23, f. 5, 21, 37, 53, 69, 81 (Df).
C. shingeni Brignoli, 1983c: 454 (lapsus).
C. s. Yaginuma, 1986a: 144, f. 77.6 (f).

mf shoshoneus Chamberlin & Ivie, 1932....................USA
C. s. Chamberlin & Ivie, 1932: 16, pl. 4, f. 37-39 (Dmf).

f signatus Keyserling, 1881....................Peru
C. s. Keyserling, 1881c: 287, pl. 11, f. 12 (Df).
C. s. Roth, 1967c: 316, pl. 51, f. 11 (f).

mf signifer Simon, 1886....................USA, Canada, Alaska
C. s. Simon, 1886d: 56 (Dmf).
C. morosus Emerton, 1923: 240, f. 5 (m, misidentified).
C. s. Chamberlin & Ivie, 1932: 12, pl. 3, f. 24-25, pl. 4, f. 32 (mf).
C. s. Roth, 1952b: 207, pl. 2, f. 16, 32 (mf).
C. s. Forster, 1970b: 13, f. 8.
C. s. Bennett, 1992: 4, f. 4-6 (f).

mf simplex Roth, 1952....................USA
C. s. Roth, 1952b: 216, pl. 1, f. 9-12 (Dmf).

f sinuosus Fox, 1937....................Canada
C. s. Fox, 1937c: 174, f. 1-2 (Df).

f strandi Kolosváry, 1934....................Transylvania
C. s. Kolosváry, 1934c: 43, f. 7 (Df).

f striatipes Bösenberg & Strand, 1906....................Japan
C. s. Bösenberg & Strand, 1906: 295, f. 5 (Df).
C. s. Ono, 1981d: 39, f. 1-5 (f).

mf tardatus (Chamberlin, 1919)....................USA
Parauximus t. Chamberlin, 1919b: 3, pl. 1, f. 2 (Dm).
Namopsilus pletus Chamberlin, 1919b: >


‹bertragung unterbrochen

. t. Chamberlin, 1924b: 669.
C. t. Chamberlin & Ivie, 1932: 23, pl. 6, f. 56 (f).

mf tetricus (C. L. Koch, 1839) *....................Europe
Amaurobius t. C. L. Koch, 1839a: 43, f. 462 (Df).
Clubiona t. Walckenaer, 1842: 482.
Amaurobius t. L. Koch, 1855: 164, f. 3 (f, Dm).
C. t. L. Koch, 1868: 50, pl. 2, f. 24-25 (mf).
C. t. Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 8 (mf).
C. t. Chamberlin & Ivie, 1932: 11, pl. 3, f. 22-23 (m).
C. t. Dahl, 1931: 9, f. 11-15 (mf).
C. t. Lehtinen, 1967: 450, f. 267, 272 (mf).
C. t. Loksa, 1969: 116, f. 79B, D, H (mf).
C. t. Brignoli, 1977f: 31, f. 5, 8 (m).
C. t. Maurer & Thaler, 1988:336, f. 3-5, 9 (m).
C. t. Heimer & Nentwig, 1991: 360, f. 935 (mf).
C. t. Maurer, 1992: 157, f. 18, 23-24 (f).
C. t. Grothendieck & Kraus, 1994: 261, f. 2a-b (m).

mf tius Chamberlin & Ivie, 1932....................USA
C. reticulatus t. Chamberlin & Ivie, 1932: 19 (Dmf).
C. t. Crawford, 1988: 28 (elevated from subspecies).

mf tottoriensis Ihara, 1994....................Japan
C. t. Ihara, 1994: 90, f. 10-16 (Dmf).

mf triangulus Paik, 1966....................Korea
C. t. Paik, 1966b: 34, f. 3, 10-16 (Dmf).
C. t. Paik, 1978e: 309, f. 134.1-6 (mf).

mf tsurusakii Ihara, 1993....................Japan
C. t. Ihara, 1993: 121, f. 15-18 (Dmf).

mf vignai Brignoli, 1977....................France, Italy
C. v. Brignoli, 1977f: 32, f. 11 (Df).
C. v. Maurer & Thaler, 1988: 334, f. 6-8, 10 (Dm).

mf yoshiakii Yaginuma, 1968....................Japan
C. y. Yaginuma, 1968a: 31, f. 1-8 (Dmf).
C. y. Yaginuma, 1986a: 145, f. 78.7 (mf).

Gen. Dolichocybaeus Kishida, 1968

D. Kishida, in Komatsu, 1968b: 16, type "Dolichocybaeus" ishikawai Kishida, in Komatsu, 1961 (the type species was originally described in a generic nomen nudum).

f anaiwaensis Komatsu, 1968....................Japan
D. a. Komatsu, 1968b: 19, f. 16, 32, 48, 64, 75, 93 (Df).
D. anaiwensis Brignoli, 1983c: 485 (lapsus).
Cybaeus a. Yaginuma, 1986a: 145, f. 77.11 (f).

f ashikitaensis Komatsu, 1968....................Japan
D. a. Komatsu, 1968b: 20, f. 14, 30, 46, 62, 73, 78, 95 (Df).
Cybaeus a. Yaginuma, 1986a: 145, f. 77.9 (f).

f fuujinensis Komatsu, 1968....................Japan
D. f. Komatsu, 1968b: 21, f. 15, 31, 47, 63, 94 (Df).
D. fujinensis Brignoli, 1983c: 455 (lapsus).

mf gotoensis Yamaguchi & Yaginuma, 1971....................Japan
D. g. Yamaguchi & Yaginuma, 1971: 176, f. 5-7 (Dmf).
Cybaeus g. Yaginuma, 1986a: 145, f. 78.5 (mf).

f ishikawai Kishida, 1961 *....................Japan
D. isikawai Kishida, in Komatsu, 1961a: 32, f. 20A-E (Df; N.B.: spelling ishikawai attributed to Kishida).
D. i. Komatsu, 1968b: 17, f. 10, 26, 42, 58, 90 (f).
Cybaeus i. Yaginuma, 1986a: 145, f. 77.3 (f).

mf kirigaminensis (Komatsu, 1963)....................Japan
Cybaeus k. Komatsu, 1963a: 7, pl. IV, f. A-H (Dmf).
D. k. Komatsu, 1968b: 20, f. 13, 29, 45, 61, 96 (Tf from Cybaeus).
Cybaeus k. Yaginuma, 1986a: 145, f. 77.1 (mf).

f kompiraensis Komatsu, 1968....................Japan
D. k. Komatsu, 1968b: 18, f. 12, 28, 44, 60, 92 (Df).

mf kuramotoi (Yaginuma, 1963)....................Japan
Cybaeus k. Yaginuma, 1963b: 54, 57, f. g-h (Dmf).
D. k. Yaginuma, 1970a: 245, f. 10 (Tf from Cybaeus).
Cybaeus k. Yaginuma, 1986a: 145, f. 78.4 (mf).

f nichikoensis Komatsu, 1968....................Japan
D. n. Komatsu, 1968a: 37, f. 7-12 (Df).

f nishikawai Komatsu, 1968....................Japan
D. n. Komatsu, 1968b: 18, f. 11, 27, 43, 59, 74, 76, 91 (Df).
Cybaeus n. Yaginuma, 1986a: 145, f. 77.10 (f).

mf okafujii (Yaginuma, 1963)....................Japan
Cybaeus o. Yaginuma, 1963b: 54, 58, f. e-f (Dmf).
D. o. Yaginuma, 1970d: 663 (Tmf from Cybaeus).
Cybaeus o. Yaginuma, 1986a: 145, f. 78.3 (mf).

f shinkaii Komatsu, 1970....................Japan
D. s. Komatsu, 1970b: 13, f. 4 (Df).

f takasawaensis Komatsu, 1970....................Japan
D. t. Komatsu, 1970b: 15, f. 1-3 (Df).

f uenoi Yaginuma, 1970....................Japan
D. u. Yaginuma, 1970a: 244, f. 6-9, 11-12 (Df).
D. u. Yaginuma, Yamaguchi & Nishikawa, 1976: 828, f. 16-21 (f).
Cybaeus u. Yaginuma, 1986a: 145, f. 78.1 (f).

mf whanseunensis (Paik & Namkung, 1967)....................Korea
Cybaeus w. Paik & Namkung, 1967: 23, f. 6-13 (Dmf).
D. w. Paik, Yaginuma & Namkung, 1969: 824 (Tmf from Cybaeus).
D. w. Paik, 1978e: 311, f. 135.1-10 (mf).

Gen. Heterocybaeus Komatsu, 1968

H. Komatsu, 1968b: 13, type H. ryusenensis Komatsu, 1968.

f akaanaensis Komatsu, 1968....................Japan
H. a. Komatsu, 1968b: 16, f. 17, 33, 49, 65, 86 (Df).

f ryusenensis Komatsu, 1968 *....................Japan
H. r. Komatsu, 1968b: 13, f. 18, 34, 50, 66, 72, 80, 87 (Df).
Cybaeus r. Yaginuma, 1986a: 145, f. 77.8 (f).

f senzokuensis Komatsu, 1968....................Japan
H. s. Komatsu, 1968b: 14, f. 19, 35, 51, 67, 71, 88 (Df).
Cybaeus s. Yaginuma, 1986a: 145, f. 77.5 (f).

f zenifukiensis Komatsu, 1968....................Japan
H. z. Komatsu, 1968b: 15, f. 20, 36, 52, 68, 89 (Df).

Gen. Symposia Simon, 1898

N.B.: placed in Amaurobioidea incertae sedis by Lehtinen, 1967: 266 and here by Brignoli, 1983c: 486.

mf bifurca Roth, 1967....................Venezuela
S. b. Roth, 1967c: 339, pl. 53, f. 9-10 (Dmf).

mf columbiana Müller & Heimer, 1988....................Colombia
S. c. Müller & Heimer, 1988a: 209, f. 1-10 (Dmf).

f dubiosa Roth, 1967....................Venezuela
S. d. Roth, 1967c: 340, pl. 53, f. 11 (Df).

mf sexoculata Roth, 1967....................Venezuela
S. s. Roth, 1967c: 337, pl. 53, f. 5-7 (Dmf).

mf silvicola Simon, 1898 *....................Venezuela
S. s. Simon, 1898g: 90 (Dmf).
S. s. Roth, 1967c: 336, pl. 53, f. 1-4 (mf).
S. s. Lehtinen, 1967: 438, f. 93, 95 (mf).

mf umbrosa Simon, 1898....................Venezuela
S. u. Simon, 1898g: 90 (Dmf).
S. u. Simon, 1898a: 239, f. 237-239 (mf).
S. u. Roth, 1967c: 338, pl. 53, f. 8 (m).
S. u. Lehtinen, 1967: 438, f. 94, 97 (mf).

Gen. Vagellia Simon, 1899

N.B.: considered a junior synonym of Zoica Simon, 1896 and placed in the Zoridae by Lehtinen, 1967: 274 but not by Lehtinen & Hippa, 1979: 14, who excluded it from the Lycosidae (where they transferred Zoica); placed here by Brignoli, 1983c: 486.

f helveola Simon, 1899 *....................Sumatra
V. h. Simon, 1899a: 101 (Df).