The World Spider Catalog, Version 11.5
by Norman I. Platnick

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FAM. CYATHOLIPIDAE Simon, 1894a: 711 [urn:lsid:amnh.org:spiderfam:0026]

Last updated May 21, 2010

N.B.: considered a senior synonym of Teemenaaridae by Platnick, 1979c: 116.

Transferred to other families:
Cyatholipulus Petrunkevitch, 1930 -- see Styposis under Theridiidae.
Cyathidea Simon, 1907 -- see Wendilgarda under Theridiosomatidae.

In synonymy:
Moero O. P.-Cambridge, 1904 = Cyatholipus Simon, 1894 (Griswold, 1987e: 505, after Petrunkevitch, 1928: 233).

Homonym replaced:
Toddiana Forster, 1988 -- see Forstera.

Gen. Alaranea Griswold, 1997 [urn:lsid:amnh.org:spidergen:00625]

A. Griswold, 1997a: 67, type A. merina Griwold, 1997.

mf alba Griswold, 1997....................Madagascar [urn:lsid:amnh.org:spidersp:006447]
A. a. Griswold, 1997a: 76, f. 55, 57-58, 67-69, 78-83, 92 (Dmf).
A. a. Griswold, 2001: 44, f. 54A, 55A, 56B (mf).

mf ardua Griswold, 1997....................Madagascar [urn:lsid:amnh.org:spidersp:006448]
A. a. Griswold, 1997a: 78, f. 84-89, 93-97 (Dmf).
A. a. Griswold, 2001: 44, f. 51A-C, 54B, 55C, 56A, C, E (mf).

mf betsileo Griswold, 1997....................Madagascar [urn:lsid:amnh.org:spidersp:006449]
A. b. Griswold, 1997a: 68, f. 2-4, 6-14, 49-54, 56, 59-64, 74-75, 90 (Dmf).
A. b. Griswold, 2001: 45, f. 4C, E, H, 52A-C, 54C, 55E, 56D (mf).

mf merina Griswold, 1997 *....................Madagascar [urn:lsid:amnh.org:spidersp:006450]
A. m. Griswold, 1997a: 72, f. 5, 65-66, 70-73, 76-77, 91 (Dmf).
A. m. Griswold, 2001: 45, f. 8B, 51D, 53A-D, 55B, D, F, 56F (mf).

Gen. Buibui Griswold, 2001 [urn:lsid:amnh.org:spidergen:03390]

B. Griswold, 2001: 46, type B. claviger Griswold, 2001.

mf abyssinica Griswold, 2001....................Ethiopia [urn:lsid:amnh.org:spidersp:037021]
B. a. Griswold, 2001: 47, f. 7D-E, 13A-D, 57A-C, 58A-C, 60A-C, 68A, C, E (Dmf).

mf claviger Griswold, 2001 *....................Kenya [urn:lsid:amnh.org:spidersp:037022]
B. c. Griswold, 2001: 49, f. 59A-C, 61A-D, 64A (Dmf).

mf cyrtata Griswold, 2001....................Congo [urn:lsid:amnh.org:spidersp:037023]
B. c. Griswold, 2001: 50, f. 4D, 7C, 58D-F, 62A-D, 64B (Dmf).

mf kankamelos Griswold, 2001....................Cameroon, Bioko [urn:lsid:amnh.org:spidersp:037024]
B. k. Griswold, 2001: 51, f. 63, 64C, 65A-C (Dmf).

mf orthoskelos Griswold, 2001....................Congo [urn:lsid:amnh.org:spidersp:037025]
B. o. Griswold, 2001: 52, f. 60D-E, 64D, 66A-D (Dmf).

Gen. Cyatholipus Simon, 1894 [urn:lsid:amnh.org:spidergen:00626]

N.B.: transferred here from the Theridiosomatidae by Davies, 1978: 301 (sub Teemenaaridae) and Wunderlich, 1978b: 33; considered a senior synonym of Moero O. P.-Cambridge, 1904 by Griswold, 1987e: 505, after Petrunkevitch, 1928: 233.

Transferred to other genera:
C. dentipes Simon, 1894 -- see Pokennips.

In synonymy:
C. quadrimaculatus (O. P.-Cambridge, 1904) = C. quadrimaculatus Simon, 1894 (Griswold, 1987e: 505, after Petrunkevitch, 1928: 233).

mf avus Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006451]
C. a. Griswold, 1987e: 512, f. 24, 26, 28, 36-38, 45-47, 55 (Dmf).
C. a. Griswold, 2001: 55, f. 67B-C, 70D (mf).

f hirsutissimus Simon, 1894 *....................South Africa [urn:lsid:amnh.org:spidersp:006453]
C. h. Simon, 1894a: 712, f. 786 (Df).
C. h. Griswold, 1987e: 507, f. 1-2, 39-41 (f).

f icubatus Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006454]
C. i. Griswold, 1987e: 514, f. 5, 48-50, 56 (Df).

mf isolatus Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006455]
C. i. Griswold, 1987e: 516, f. 51-53, 57 (Df).
C. i. Griswold, 2001: 56, f. 67A, 68B, D, F, 69A-D, 70A-C (f, Dm).

mf quadrimaculatus Simon, 1894....................South Africa [urn:lsid:amnh.org:spidersp:006456]
C. q. Simon, 1894a: 713, f. 787 (Dmf).
Moero quadrimaculata O. P.-Cambridge, 1904: 157, pl. 11, f. 4 (Dmf).
C. q. Griswold, 1987e: 508, f. 3-4, 7, 20-23, 25, 27, 42-44, 54 (mf, S).
C. q. Forster, 1988: 18, f. 31, 51 (f).

m tortilis Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006457]
C. t. Griswold, 1987e: 510, f. 6, 29-35 (Dm).

Gen. Forstera Koçak & Kemal, 2008 [urn:lsid:amnh.org:spidergen:03829]

F. Koçak & Kemal, 2008a: 2, replacement name for Toddiana Forster, 1988: 31 [urn:lsid:amnh.org:spidergen:00636], type T. daviesae Forster, 1988, preoccupied in the Lepidoptera; objective senior synonym of Queenslandiana Özdikmen, 2009: 123 (superfluous replacement name).

mf daviesae (Forster, 1988) *....................Queensland [urn:lsid:amnh.org:spidersp:006478]
Toddiana d. Forster, 1988: 32, f. 27, 29, 59, 111-112, 117-122 (Dmf).
Toddiana d. Griswold, 2001: 39, f. 6C, 9C-D, 14F, 29D-F, 30D, 38A-C, 39A-D, 40A-C (mf).

Gen. Hanea Forster, 1988 [urn:lsid:amnh.org:spidergen:00627]

H. Forster, 1988: 26, type H. paturau Forster, 1988.

m paturau Forster, 1988 *....................New Zealand [urn:lsid:amnh.org:spidersp:006458]
H. p. Forster, 1988: 26, f. 46-47, 91-94 (Dm).
H. p. Griswold, 2001: 40, f. 41A-C, 42A-B (m).

Gen. Ilisoa Griswold, 1987 [urn:lsid:amnh.org:spidergen:00628]

I. Griswold, 1987e: 517, type I. knysna Griswold, 1987.

mf conjugalis Griswold, 2001....................South Africa [urn:lsid:amnh.org:spidersp:037026]
I. c. Griswold, 2001: 58, f. 71A-C, 72A-C, 73A, C-D, 75A-C (Dmf).

f hawequas Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006459]
I. h. Griswold, 1987e: 520, f. 58-59, 69-71, 114 (Df).
I. h. Griswold, 2001: 59, f. 73B, 75D (f).

m knysna Griswold, 1987 *....................South Africa [urn:lsid:amnh.org:spidersp:006460]
I. k. Griswold, 1987e: 518, f. 15, 60-68 (Dm).
I. k. Griswold, 2001: 60, f. 73E-F, 74A-C (m).

Gen. Isicabu Griswold, 1987 [urn:lsid:amnh.org:spidergen:00629]

I. Griswold, 1987e: 522, type I. reavelli Griswold, 1987.

Transferred to other genera:
I. capensis Griswold, 1987 -- see Ubacisi.

mf henriki Griswold, 2001....................Tanzania [urn:lsid:amnh.org:spidersp:037027]
I. h. Griswold, 2001: 61, f. 76, 77A, 78A-D, 79A-C, 85B, D, F (Dmf).

mf kombo Griswold, 2001....................Tanzania [urn:lsid:amnh.org:spidersp:037028]
I. k. Griswold, 2001: 63, f. 4B, F, 7F, 77B-C, 79D-F, 80, 81A-D, 82A-C, 85A, C, E (Dmf).

mf margrethae Griswold, 2001....................Tanzania [urn:lsid:amnh.org:spidersp:037029]
I. m. Griswold, 2001: 64, f. 77D, 83, 84A-D, 85G (Dmf).

mf reavelli Griswold, 1987 *....................South Africa [urn:lsid:amnh.org:spidersp:006462]
I. r. Griswold, 1987e: 523, f. 8, 14, 72-74, 84-89, 93-95, 115 (Dmf).
I. r. Griswold, 2001: 66, f. 79G-I, 86A-C (mf).

m zuluensis Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006463]
I. z. Griswold, 1987e: 525, f. 90, 96-101 (Dm).

Gen. Kubwa Griswold, 2001 [urn:lsid:amnh.org:spidergen:03401]

K. Griswold, 2001: 67, type K. singularis Griswold, 2001.

mf singularis Griswold, 2001 *....................Tanzania [urn:lsid:amnh.org:spidersp:037030]
K. s. Griswold, 2001: 67, f. 14A, 87A-C, 88A-D, 89A-C, 90A-C, 91A-C (Dmf).

Gen. Lordhowea Griswold, 2001 [urn:lsid:amnh.org:spidergen:03403]

L. Griswold, 2001: 33, type L. nesiota Griswold, 2001.

mf nesiota Griswold, 2001 *....................Lord Howe Is. [urn:lsid:amnh.org:spidersp:037031]
L. n. Griswold, 2001: 34, f. 22A-C, 23A-D, 24A-C, 25A-B (Dmf).

Gen. Matilda Forster, 1988 [urn:lsid:amnh.org:spidergen:00630]

M. Forster, 1988: 33, type M. australia Forster, 1988.

mf australia Forster, 1988 *....................Queensland, New South Wales [urn:lsid:amnh.org:spidersp:006464]
M. a. Forster, 1988: 33, f. 108, 123-140 (Dmf).
M. a. Griswold, 2001: 35, f. 14D, 26A-C, 27A-D, 28A-C, 29A-C, 30A-C (mf).

Gen. Pembatatu Griswold, 2001 [urn:lsid:amnh.org:spidergen:03410]

P. Griswold, 2001: 69, type P. embamba Griswold, 2001.

mf embamba Griswold, 2001 *....................Kenya, Tanzania [urn:lsid:amnh.org:spidersp:037032]
P. e. Griswold, 2001: 70, f. 1, 93A-D, 94A-B, 99A, C, E (Dmf).

mf gongo Griswold, 2001....................Kenya [urn:lsid:amnh.org:spidersp:037033]
P. g. Griswold, 2001: 71, f. 92B-C, 95A-D, 96B (Dmf).

mf mafuta Griswold, 2001....................Kenya [urn:lsid:amnh.org:spidersp:037034]
P. m. Griswold, 2001: 73, f. 6A-B, D-E, 7A-B, 92A, 94C, 96A, 97A-D, 98A-C (Dmf).

Gen. Pokennips Griswold, 2001 [urn:lsid:amnh.org:spidergen:03412]

P. Griswold, 2001: 74, type Cyatholipus dentipes Simon, 1894.

mf dentipes (Simon, 1894) *....................Jamaica [urn:lsid:amnh.org:spidersp:006452]
Cyatholipus d. Simon, 1894a: 713, f. 788-789 (Dm).
Cyatholipus d. Wunderlich, 1978b: 39, f. 1-4 (m, Df).
Cyatholipus d. Wunderlich, 1986: 223, f. 248 (m).
P. d. Griswold, 2001: 75, f. 5E, 99B, D, F, 100A-C, 101A-D, 102A-C, 103A-C (Tmf from Cyatholipus).

Gen. Scharffia Griswold, 1997 [urn:lsid:amnh.org:spidergen:00631]

S. Griswold, 1997b: 270, type S. chinja Griswold, 1997.

mf chinja Griswold, 1997 *....................Tanzania [urn:lsid:amnh.org:spidersp:006465]
S. c. Griswold, 1997b: 272, f. 5-23, 25, 38, 40, 44-46 (Dmf).
S. c. Griswold, 2001: 77, f. 4A, 8A, 104C-F, 105A-D, 107A-C, 108A, D-E, 109A (mf).

mf holmi Griswold, 1997....................Kenya [urn:lsid:amnh.org:spidersp:006466]
S. h. Griswold, 1997b: 277, f. 27-29, 32-36, 39 (Dmf).
S. h. Griswold, 2001: 78, f. 104A-B, 108B, 109C (mf).

mf nyasa Griswold, 1997....................Malawi [urn:lsid:amnh.org:spidersp:006467]
S. n. Griswold, 1997b: 278, f. 24, 26, 37, 41-43, 47-53 (Dmf).
S. n. Griswold, 2001: 79, f. 106A-C, 108C, 109B (mf).

m rossi Griswold, 1997....................Tanzania [urn:lsid:amnh.org:spidersp:006468]
S. r. Griswold, 1997b: 280, f. 1, 30-31, 54-57 (Dm).
S. r. Griswold, 2001: 79, f. 107D-E, 109D (m).

Gen. Teemenaarus Davies, 1978 [urn:lsid:amnh.org:spidergen:00632]

T. Davies, 1978: 294, type T. silvestris Davies, 1978.

mf silvestris Davies, 1978 *....................Queensland [urn:lsid:amnh.org:spidersp:006469]
T. s. Davies, 1978: 294, f. 1-15, 17-20 (Dmf).
T. s. Forster, 1988: 30, f. 29-30, 109-110, 113-116 (mf).
T. s. Coddington, 1990: 20, f. 107 (m).
T. s. Griswold, 2001: 36, f. 30E, 31A-C, 32A-D, 33A-C, 34A-C (mf).

Gen. Tekella Urquhart, 1894 [urn:lsid:amnh.org:spidergen:00633]

N.B. transferred from the Theridiidae to the Theridiosomatidae by Levi & Levi, 1962: 29, and here from the Theridiosomatidae by Davies, 1978: 301 and Wunderlich, 1978b: 33.

mf absidata Urquhart, 1894....................New Zealand [urn:lsid:amnh.org:spidersp:006470]
T. a. Urquhart, 1894: 211 (Df).
T. a. Forster, 1988: 19, f. 11-26, 32-33, 36-37, 58, 60-67 (f, Dm, removed from S of T. nemoralis, contra Parrott, 1946: 71).
T. a. Griswold et al., 1998: 68, f. 17A-B (m).
T. a. Griswold, 2001: 41, f. 14E, 43A, C, E, 44A-C, 45A-D, 46A-C, 47A (mf).

mf bisetosa Forster, 1988....................New Zealand [urn:lsid:amnh.org:spidersp:006471]
T. b. Forster, 1988: 25, f. 42-43, 53, 86-90 (Dmf).

mf lineata Forster, 1988....................New Zealand [urn:lsid:amnh.org:spidersp:006472]
T. l. Forster, 1988: 22, f. 7-8, 44-45, 49, 57, 74-79 (Dmf).

mf nemoralis (Urquhart, 1889) *....................New Zealand [urn:lsid:amnh.org:spidersp:006473]
Linyphia n. Urquhart, 1889a: 140 (Dmf).
T. n. Bryant, 1933b: 19, pl. 2, f. 12-13, 17 (mf).
T. n. Wunderlich, 1978b: 39, f. 15-23 (mf).
T. n. Wunderlich, 1986: 223, f. 247 (m).
T. n. Forster, 1988: 21, f. 38-39, 48, 54, 68-73 (mf, not a senior S of T. absidata).

mf unisetosa Forster, 1988....................New Zealand [urn:lsid:amnh.org:spidersp:006474]
T. u. Forster, 1988: 23, f. 40-41, 50, 52, 80-85 (Dmf).

Gen. Tekellatus Wunderlich, 1978 [urn:lsid:amnh.org:spidergen:00634]

T. Wunderlich, 1978b: 38, type T. lamingtoniensis Wunderlich, 1978.

m lamingtoniensis Wunderlich, 1978 *....................Queensland [urn:lsid:amnh.org:spidersp:006475]
T. l. Wunderlich, 1978b: 39, f. 5-14 (Dm).
T. l. Wunderlich, 1986: 222, f. 245-246 (m).
T. l. Griswold, 2001: 38, f. 35, 36A-D, 37A-B (m).

Gen. Tekelloides Forster, 1988 [urn:lsid:amnh.org:spidergen:00635]

T. Forster, 1988: 26, type T. australis Forster, 1988.

mf australis Forster, 1988 *....................New Zealand [urn:lsid:amnh.org:spidersp:006476]
T. a. Forster, 1988: 28, f. 3-4, 9-10, 28, 95-103 (Dmf).
T. a. Griswold, 2001: 42, f. 9A-B, 14B, 43B, D, F, 47B-D, 48A-C, 49A-D, 50A-C (mf).

mf flavonotatus (Urquhart, 1891)....................New Zealand [urn:lsid:amnh.org:spidersp:006477]
Ariamnes flavo-notatus Urquhart, 1891a: 136 (Df).
T. f. Forster, 1988: 28, f. 34-35, 55-56, 104-107 (Tf from Ariamnes=Argyrodes, Dm).
T. f. Paquin, Vink & Dupérré, 2010: 58, f. 32.2-3 (m).

Gen. Ubacisi Griswold, 2001 [urn:lsid:amnh.org:spidergen:03424]

U. Griswold, 2001: 79, type Isicabu capensis Griswold, 1987.

mf capensis (Griswold, 1987) *....................South Africa [urn:lsid:amnh.org:spidersp:006461]
Isicabu c. Griswold, 1987e: 528, f. 75-83, 91-92, 102-110, 116 (Dmf).
U. c. Griswold, 2001: 80, f. 110, 111A-D, 112A-C, 113A-C, 114A-C (Tmf from Isicabu).

Gen. Ulwembua Griswold, 1987 [urn:lsid:amnh.org:spidergen:00637]

U. Griswold, 1987e: 532, type U. pulchra Griswold, 1987.

mf antsiranana Griswold, 1997....................Madagascar [urn:lsid:amnh.org:spidersp:006479]
U. a. Griswold, 1997a: 59, f. 1, 16-18, 22-26, 35, 38 (Dmf).
U. a. Griswold, 2001: 82, f. 115, 116A, 125A, 133B (mf).

mf denticulata Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006480]
U. d. Griswold, 1987e: 536, f. 112, 121-122, 134-136 (Df).
U. d. Griswold, 2001: 82, f. 15A-C, 117, 118A-D, 119A-C, 120B-C, 133C (f, Dm).

mf nigra Griswold, 2001....................Madagascar [urn:lsid:amnh.org:spidersp:037035]
U. n. Griswold, 2001: 84, f. 121A-B, 122A-D, 123A-C, 133D (Dmf).

mf outeniqua Griswold, 1987....................South Africa [urn:lsid:amnh.org:spidersp:006481]
U. o. Griswold, 1987e: 538, f. 9-13, 113, 117-120, 126-127, 137-139, 143-151 (Dmf).
U. o. Griswold, 2001: 85, f. 12A-C, 121C, 124A-D, 125D-F, 133A (mf).

mf pulchra Griswold, 1987 *....................South Africa [urn:lsid:amnh.org:spidersp:006482]
U. p. Griswold, 1987e: 534, f. 111, 123-125, 128-133, 140-142 (Dmf).
U. p. Griswold, 2001: 86, f. 120A, 126, 127A-D, 128A-C (mf).

mf ranomafana Griswold, 1997....................Madagascar [urn:lsid:amnh.org:spidersp:006483]
U. r. Griswold, 1997a: 63, f. 15, 19-21, 27-28, 31-34, 36, 39 (Dmf).
U. r. Griswold, 2001: 86, f. 16A, C, E, 116B, 125B, 129A-C (mf).
U. r. Jocqué & Dippenaar-Schoeman, 2006: 102, f. 28b-f (mf).

mf usambara Griswold, 2001....................Tanzania [urn:lsid:amnh.org:spidersp:037036]
U. u. Griswold, 2001: 87, f. 125C, 130A-C, 131A-D, 132A-C, 133E (Dmf).

Gen. Umwani Griswold, 2001 [urn:lsid:amnh.org:spidergen:03425]

U. Griswold, 2001: 88, type U. artigamos Griswold, 2001.

m anymphos Griswold, 2001....................Malawi [urn:lsid:amnh.org:spidersp:037037]
U. a. Griswold, 2001: 89, f. 134D-F, 135A-C, 136A-B (Dm).

mf artigamos Griswold, 2001 *....................Tanzania [urn:lsid:amnh.org:spidersp:037038]
U. a. Griswold, 2001: 90, f. 134A-C, 136C-E, 137A-D, 138B, D, F (Dmf).

Gen. Uvik Griswold, 2001 [urn:lsid:amnh.org:spidergen:03426]

U. Griswold, 2001: 91, type U. vulgaris Griswold, 2001.

mf vulgaris Griswold, 2001 *....................Congo, Uganda [urn:lsid:amnh.org:spidersp:037039]
U. v. Griswold, 2001: 92, f. 10A-D, 11A-D, 14C, 138A, C, E, 139A-C, 140A-D, 141A-C, 142A-C (Dmf).

Gen. Vazaha Griswold, 1997 [urn:lsid:amnh.org:spidergen:00638]

V. Griswold, 1997a: 65, type V. toamasina Griswold, 1997.

mf toamasina Griswold, 1997 *....................Madagascar [urn:lsid:amnh.org:spidersp:006484]
V. t. Griswold, 1997a: 65, f. 29-30, 37, 40-48 (Dmf).
V. t. Griswold, 2001: 94, f. 133F-G, 143, 144A-D, 145A-C, 146A-C (mf).

Gen. Wanzia Griswold, 1998 [urn:lsid:amnh.org:spidergen:00639]

W. Griswold, 1998a: 123, type W. fako Griswold, 1998.

mf fako Griswold, 1998 *....................Cameroon, Bioko [urn:lsid:amnh.org:spidersp:006485]
W. f. Griswold, 1998a: 125, f. 1-17 (Dmf).
W. f. Griswold, 2001: 96, f. 4G, 5A-D, 16B, D, F, 147, 148A-D, 149A-C (mf).