The Ants of Africa
SUBFAMILY DORYLINAE - Subgenus Anomma comparative study and revision 2002, amended 2007
Subgenus Anomma introduction

This page deals with the sources of information on the subgenus Anomma and gives a suggested separation of the described species into species-groupings - a key to separate the species is linked at Subgenus Anomma key to species; also examined are the Subgenus Anomma comparison of the gastral apices.

Note - At the time of writing, December 2007, Caspar Schöning and Daniel Kronauer, are preparing a fresh analysis of Dorylus taxonomy. I anticipate that may show some of my interpretations of the species to be incorrect.


Species known only from sexual stages

Dorylus (Anomma) bishyiganus van Boven - queen
Dorylus (Anomma) funereus Emery - male; junior synonyms acherontus Santschi, pardus Santschi, stygis Santschi and zumpti Santschi
Dorylus (Anomma) niarembensis van Boven - queen
Dorylus (Anomma) nigricans Illiger - male (but see below)
Dorylus (Anomma) stanleyi Forel - male

The overall size, TL ca 19 mm, and mandible shape would appear to place Dorylus atratus Smith in subgenus Alaopone and not Anomma as given by Emery (1895j: 723).


I have tried to separate what is known of the males on MALES.


Time line of Anomma forms

Name First author Present status grouping & images
nigricans - male only; workers by later deduction Illiger, 1802 Dorylus nigricans male & possible workers, Schöning, Ivory Coast
burmeisteri - worker only Shuckard, 1840 Dorylus burmeisteri yes - Humle 13, Cameroon 11
arcens - worker only Westwood, 1847b Dorylus arcens yes, Cameroun, Benin & Guinea - Humle 07
rubellus - worker only Savage, 1849 Dorylus rubellus yes, Schöning CAR
molestus - worker only Gerstæcker, 1858 Dorylus molestus yes, Schöning Kenya
erraticus - minor worker only F Smith, 1865 Dorylus erraticus no merit
pubescens j. syn. of arcens Roger, 1861a; syn. Emery, 1895j, after examining type Dorylus arcens -
funereus - male only Emery, 1895j Dorylus funereus yes, male, CAR
gerstaeckeri - worker only Emery, 1895j Dorylus gerstaeckeri yes, Cameroon 37
emeryi - worker only Mayr, 1896 Dorylus emeryi yes, Schöning, Ivory Coast; drawing CRIN Nigeria
sjoestedti (as ssp of nigricans) - worker & queen Emery, 1899e Dorylus sjoestedti yes, CAR, Degallier; Schöning, Ivory Coast
wilverthi - all forms known Emery, 1899e Dorylus wilverthi yes, Uganda 0310103 (Schöning - Zaire)
pulsi ssp of emeryi - worker only Forel, 1904d Dorylus emeryi -
kohli - worker only Wasmann, 1904b Dorylus kohli yes, Guinea - Humle 11, Humle 2
opacus (as ssp of emeryi) - worker & male known Forel, 1909b Dorylus opacus yes, Schöning, Nigeria
stanleyi - male only Forel, 1909b Dorylus stanleyi -
congolensis (as variety of kohli) - worker only Santschi, 1910c Dorylus congolensis yes, Gabon - Pongara
minor variety of kohli Santschi, 1911g Dorylus kohli -
nigritarsis queen j syn of wilverthi Strand 1911 Dorylus wilverthi -
hybridus [hybrida] variety of burmeisteri Santschi, 1912b Dorylus burmeisteri -
ornatus (as variety of burmeisteri) - worker only Santschi, 1912b Dorylus ornatus yes, Uganda III-138 & Tanzania Mahale 05-03-02, 02-03-02, 08-03-02
pallidus variety of burmeisteri Santschi, 1912b Dorylus burmeisteri -
mayri - worker only Santschi, 1912b Dorylus mayri yes, Uganda III-135
quadratus (as variety of gerstaeckeri) - worker only Santschi, 1914a Dorylus quadratus yes, Tanzania, Mahale 03-03-02
rufescens (as stirps of nigricans) - worker & queen Santschi, 1915c - redefined from Wasmann, 1904b Dorylus rufescens yes, CAR, site GJ
frenisyi ssp of kohli Forel, 1916 Dorylus kohli -
sjoestedtiwilverthi j syn of wilverthi Wasmann, 1917 Dorylus wilverthi -
victoriae (as var. of kohli) - worker only Santschi, 1921c Dorylus victoriae yes, Ivory Coast - Humle 12; Guinea - Humle 3; Ghana 06G0066; Schöning, Ivory Coast labelled D. mayri
chapini ssp of kohli Wheeler, 1922 Dorylus kohli -
langi j syn of victoriae Wheeler, 1922 Dorylus victoriae -
militaris (as var. of kohli) - worker only Santschi, 1923e Dorylus militaris yes, Guinea - Humle 06, Humle 08
terrificus (as variety of nigricans) - worker only Santschi, 1923e Dorylus terrificus yes, Guinea - Humle 09
titan - worker only Santschi, 1923e Dorylus titan yes, Guinea - Humle 04
indocilis (as var. of kohli) - worker only Santschi, 1933b Dorylus indocilis yes, Tanzania - Mahale 28-02-02
nomadas queen j syn of wilverthi Santschi, 1933b Dorylus wilverthi -
vinalli var. of titan Santschi, 1933b Dorylus titan -
acherontus - variety of funereus - male only Santschi, 1937b Dorylus funereus -
pardus - variety of funereus - male only Santschi, 1937b Dorylus funereus -
stygis - variety of funereus - male only Santschi, 1937b Dorylus funereus -
zumpti - variety of funereus - male only Santschi, 1937b Dorylus funereus -
lamottei - worker only Bernard, 1953b Dorylus lamottei yes, Guinea - Humle 05; Schöning, Ivory Coast casent0172626 labelled D. gribodoi
bishyiganus - queen only van Boven, 1972 Dorylus bishyiganus no
niarembensis - queen only van Boven, 1972 Dorylus niarembensis no
ueharae - worker only new species - this website Dorylus ueharae yes, Tanzania - Mahale 27-02-02

New key to species-groups 2007 - grouping derived from original descriptions and relationships

See also the Subgenus Anomma comparison of the gastral apices

. Major or soldier Minima Grouping
. Long-legged with petioles longer than they are wide .
1 {Dorylus burmeisteri face}Head widest mid-length, near parallel sided forward of that point, with rounded posterior angles; mandibles moderate; scapes relatively long & slender; in profile head depth ca 0.75 of HW and HL; TL max ca 12.5 mm

{Dorylus burmeisteri minima}

burmeisteri-group
burmeisteri, molestus, nigricans & rubellus
2 {Dorylus arcens face}Large, TL ca 13 mm; head widest at about the one-fifth point, with smoothly convex sides and sharp posterior angles; in profile head bulbous much deeper than other groups; mandibles very long and sharp; scapes relatively long & slender; generally more robust than burmeisteri-gp. Apart from the elongated processes at the postero-lateral extremities of the head wilverthi generally fits with the other arcens-group species {Dorylus arcens minima} arcens-group
arcens, mayri, terrificus & wilverthi
3 {Dorylus sjoestedti major}Characterised by the head being widest at the anterior end and narrowing significantly from front to back, with almost straight sides and sharp posterior angles; the latter never produced into distinct teeth (unlike wilverthi) {Dorylus sjoestedti minima} sjoestedti-group
rufescens, sjoestedti & victoriae
. Short-legged with petioles shorter than they are wide .
4 {Dorylus emery major}Relatively large, TL to 13 mm; head near square, rectangular; characterised by the short domed petiole, which seen from above is straight-sided and near square {Dorylus titan minima} emeryi-group
emeryi, opacus & titan
5 {Dorylus lamottei}Small less than 8 mm; head square, petiole short and domed/globular - possible link to Dorylus (Dorylus) {Dorylus lamottei minima} lamottei-group
lamottei & ueharae
6 {Dorylus congolensis major}Head near rectangular distinctly longer than wide and narrower posteriorly; petiole short in profile generally domed (some with a flat top); generally not more than TL 10 mm (congolensis to 12.5 mm) {Dorylus congolensis minima} kohli-group
congolensis, kohli & militaris
7 {Dorylus quadratus major}Small, TL ca 8 mm-10 mm; head fairly rectangular but narrowing somewhat after the mid-point; with quite long mandibles and legs {Dorylus quadratus minima} gerstaeckeri-group
gerstaeckeri, indocilis & quadratus

Contents Subgenus Anomma introduction
© 2007 - Brian Taylor CBiol FIBiol FRES
11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K.

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