| The Ants of Africa SUBFAMILY DORYLINAE - Subgenus Anomma comparative study and revision 2002, amended 2007 |
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| Subgenus Anomma introduction |
This page deals with the sources of information on the subgenus Anomma and gives a suggested separation of the described species into species-groupings - a key to separate the species is linked at Subgenus Anomma key to species; also examined are the Subgenus Anomma comparison of the gastral apices.
Note - At the time of writing, December 2007, Caspar Schöning and Daniel Kronauer, are preparing a fresh analysis of Dorylus taxonomy. I anticipate that may show some of my interpretations of the species to be incorrect.
Species known only from sexual stages
Dorylus
(Anomma) bishyiganus van Boven - queen
Dorylus
(Anomma) funereus Emery - male; junior synonyms acherontus
Santschi, pardus Santschi, stygis Santschi and zumpti
Santschi
Dorylus
(Anomma) niarembensis van Boven - queen
Dorylus
(Anomma) nigricans Illiger - male (but see below)
Dorylus
(Anomma) stanleyi Forel - male
The overall size, TL ca 19 mm, and mandible shape would appear to place Dorylus atratus Smith in subgenus Alaopone and not Anomma as given by Emery (1895j: 723).
I have tried to separate what is known of the males on MALES.
Time line of Anomma forms
| Name | First author | Present status | grouping & images |
| nigricans - male only; workers by later deduction | Illiger, 1802 | Dorylus nigricans | male & possible workers, Schöning, Ivory Coast |
| burmeisteri - worker only | Shuckard, 1840 | Dorylus burmeisteri | yes - Humle 13, Cameroon 11 |
| arcens - worker only | Westwood, 1847b | Dorylus arcens | yes, Cameroun, Benin & Guinea - Humle 07 |
| rubellus - worker only | Savage, 1849 | Dorylus rubellus | yes, Schöning CAR |
| molestus - worker only | Gerstæcker, 1858 | Dorylus molestus | yes, Schöning Kenya |
| erraticus - minor worker only | F Smith, 1865 | Dorylus erraticus | no merit |
| pubescens j. syn. of arcens | Roger, 1861a; syn. Emery, 1895j, after examining type | Dorylus arcens | - |
| funereus - male only | Emery, 1895j | Dorylus funereus | yes, male, CAR |
| gerstaeckeri - worker only | Emery, 1895j | Dorylus gerstaeckeri | yes, Cameroon 37 |
| emeryi - worker only | Mayr, 1896 | Dorylus emeryi | yes, Schöning, Ivory Coast; drawing CRIN Nigeria |
| sjoestedti (as ssp of nigricans) - worker & queen | Emery, 1899e | Dorylus sjoestedti | yes, CAR, Degallier; Schöning, Ivory Coast |
| wilverthi - all forms known | Emery, 1899e | Dorylus wilverthi | yes, Uganda 0310103 (Schöning - Zaire) |
| pulsi ssp of emeryi - worker only | Forel, 1904d | Dorylus emeryi | - |
| kohli - worker only | Wasmann, 1904b | Dorylus kohli | yes, Guinea - Humle 11, Humle 2 |
| opacus (as ssp of emeryi) - worker & male known | Forel, 1909b | Dorylus opacus | yes, Schöning, Nigeria |
| stanleyi - male only | Forel, 1909b | Dorylus stanleyi | - |
| congolensis (as variety of kohli) - worker only | Santschi, 1910c | Dorylus congolensis | yes, Gabon - Pongara |
| minor variety of kohli | Santschi, 1911g | Dorylus kohli | - |
| nigritarsis queen j syn of wilverthi | Strand 1911 | Dorylus wilverthi | - |
| hybridus [hybrida] variety of burmeisteri | Santschi, 1912b | Dorylus burmeisteri | - |
| ornatus (as variety of burmeisteri) - worker only | Santschi, 1912b | Dorylus ornatus | yes, Uganda III-138 & Tanzania Mahale 05-03-02, 02-03-02, 08-03-02 |
| pallidus variety of burmeisteri | Santschi, 1912b | Dorylus burmeisteri | - |
| mayri - worker only | Santschi, 1912b | Dorylus mayri | yes, Uganda III-135 |
| quadratus (as variety of gerstaeckeri) - worker only | Santschi, 1914a | Dorylus quadratus | yes, Tanzania, Mahale 03-03-02 |
| rufescens (as stirps of nigricans) - worker & queen | Santschi, 1915c - redefined from Wasmann, 1904b | Dorylus rufescens | yes, CAR, site GJ |
| frenisyi ssp of kohli | Forel, 1916 | Dorylus kohli | - |
| sjoestedtiwilverthi j syn of wilverthi | Wasmann, 1917 | Dorylus wilverthi | - |
| victoriae (as var. of kohli) - worker only | Santschi, 1921c | Dorylus victoriae | yes, Ivory Coast - Humle 12; Guinea - Humle 3; Ghana 06G0066; Schöning, Ivory Coast labelled D. mayri |
| chapini ssp of kohli | Wheeler, 1922 | Dorylus kohli | - |
| langi j syn of victoriae | Wheeler, 1922 | Dorylus victoriae | - |
| militaris (as var. of kohli) - worker only | Santschi, 1923e | Dorylus militaris | yes, Guinea - Humle 06, Humle 08 |
| terrificus (as variety of nigricans) - worker only | Santschi, 1923e | Dorylus terrificus | yes, Guinea - Humle 09 |
| titan - worker only | Santschi, 1923e | Dorylus titan | yes, Guinea - Humle 04 |
| indocilis (as var. of kohli) - worker only | Santschi, 1933b | Dorylus indocilis | yes, Tanzania - Mahale 28-02-02 |
| nomadas queen j syn of wilverthi | Santschi, 1933b | Dorylus wilverthi | - |
| vinalli var. of titan | Santschi, 1933b | Dorylus titan | - |
| acherontus - variety of funereus - male only | Santschi, 1937b | Dorylus funereus | - |
| pardus - variety of funereus - male only | Santschi, 1937b | Dorylus funereus | - |
| stygis - variety of funereus - male only | Santschi, 1937b | Dorylus funereus | - |
| zumpti - variety of funereus - male only | Santschi, 1937b | Dorylus funereus | - |
| lamottei - worker only | Bernard, 1953b | Dorylus lamottei | yes, Guinea - Humle 05; Schöning, Ivory Coast casent0172626 labelled D. gribodoi |
| bishyiganus - queen only | van Boven, 1972 | Dorylus bishyiganus | no |
| niarembensis - queen only | van Boven, 1972 | Dorylus niarembensis | no |
| ueharae - worker only | new species - this website | Dorylus ueharae | yes, Tanzania - Mahale 27-02-02 |
New key to species-groups 2007 - grouping derived from original descriptions and relationships
See also the Subgenus Anomma comparison of the gastral apices
| . | Major or soldier | Minima | Grouping |
| . | Long-legged with petioles longer than they are wide | . | |
| 1 | Head
widest mid-length, near parallel sided forward of that point, with
rounded posterior angles; mandibles moderate; scapes relatively long &
slender; in profile head depth ca 0.75 of HW and HL; TL max ca 12.5 mm |
|
burmeisteri-group burmeisteri, molestus, nigricans & rubellus |
| 2 | Large,
TL ca 13 mm; head widest at about the one-fifth point, with smoothly
convex sides and sharp posterior angles; in profile head bulbous much
deeper than other groups; mandibles very long and sharp; scapes
relatively long & slender; generally more robust than burmeisteri-gp.
Apart from the elongated processes at the postero-lateral extremities of
the head wilverthi generally fits with the other arcens-group
species |
![]() |
arcens-group arcens, mayri, terrificus & wilverthi |
| 3 | Characterised
by the head being widest at the anterior end and narrowing significantly
from front to back, with almost straight sides and sharp posterior
angles; the latter never produced into distinct teeth (unlike wilverthi) |
![]() |
sjoestedti-group rufescens, sjoestedti & victoriae |
| . | Short-legged with petioles shorter than they are wide | . | |
| 4 | Relatively
large, TL to 13 mm; head near square, rectangular; characterised by the
short domed petiole, which seen from above is straight-sided and near
square |
![]() |
emeryi-group emeryi, opacus & titan |
| 5 | Small
less than 8 mm; head square, petiole short and domed/globular - possible
link to Dorylus (Dorylus) |
![]() |
lamottei-group lamottei & ueharae |
| 6 | Head
near rectangular distinctly longer than wide and narrower posteriorly;
petiole short in profile generally domed (some with a flat top);
generally not more than TL 10 mm (congolensis to 12.5 mm) |
![]() |
kohli-group congolensis, kohli & militaris |
| 7 | Small,
TL ca 8 mm-10 mm; head fairly rectangular but narrowing somewhat after
the mid-point; with quite long mandibles and legs |
![]() |
gerstaeckeri-group gerstaeckeri, indocilis & quadratus |
| Subgenus Anomma introduction | © 2007 - Brian Taylor CBiol
FIBiol FRES 11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K. |
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