The Ants of Africa
Genus Camponotus subgenus Tanaemyrmex
Camponotus maculatus varieties
Camponotus maculatus main entry
{Camponotus maculatus type form}

Camponotus (Tanaemyrmex) maculatus (Fabricius)

return to key {link to the Hymenoptera Name Server} Type location West Africa (Fabricius, 1782: 491, worker; Mayr, 1862: 654, queen & male; Donisthorpe, 1915a: 221, redescription of the type; Wheeler, 1922, reported the location as "in Africa aequinoctali" adding probably Sierra Leone) .

The original Fabricius (1793) description is at {original description}. Mayr (1862: 654) gave a further description which is at {original description}.

C. maculatus sensu stricto

{Camponotus (Tanaemyrmex) maculatus} Major worker - TL 14.0, HL 3.99, HW 3.86, SL 3.29, PW 2.22
Minor worker - TL 9.50, HL 2.24, HW 1.31, SL 3.11, PW 1.37
Colour orange to dark red-brown, darkest on gaster but characteristically lateral light patches on tergites 1 to 3, shiny. Erect coarse hairs long but sparse, and very sparse pilosity. Declivity of propodeum smoothly rounded. Petiole a rounded node in the minor but sharper in the major.

The foregoing was my description of the form common at CRIN and shown in my drawing. The specimens sent to me from Cameroon (see below) match this exactly. Wheeler (1922) referred to the description by Donisthorpe of the "Fabrician type in the Banks Collection" and noted that specimens collected by Lang & Chapin at several Congo locations "agree perfectly" but "have a few short erect hairs on the gular surface of the head". I do not recall seeing any gular hairs on the CRIN specimens and there are none on the Cameroon examples. I have to conclude that what I drew and the Cameroon specimens represent the C. maculatus sensu stricto and that specimens with gular hairs are of a related species.

I have transcribed Donisthorpe's description or rather re-description of the type specimen on the linked page - Camponotus maculatus and the Cameroun specimens are a perfect match in all aspects.

{Camponotus maculatus type minor}Minor

The photomontages of what I regard as the type form are of specimens collected in Cameroun - south-western tropical coastal forest area between Edéa and Campo (McKey Wolbachia project) - Cameroon 99 from location Kribi, 15 April 2001, in herbaceous vegetation about 20 cm tall, in garden of the Catholic Mission.

General morphology

{Camponotus maculatus varieties}A very widely distributed species with apparently great variability. The name derives from the French "maculé" or spotted. Forel (1879) suggested the colour pattern on the gaster could be either yellow with dark patches or the converse (variegatus); he reported locations as "Sénégal, Cap Verd, Guineé, Egypte, Inde (coll. Sauss)". That probably would place the type location as Cap Vert, near Dakar, Senegal (as more precisely stated for collections by Sauss of Camp. foraminosus). For cognata (from Southern Africa) he noted it as the race which held the largest , major TL up to 16 mm, females to 18 mm plus, and most highly sculptured (with puncturate rugo-reticulation on the head and thorax; plus transverse rugae on the gaster) forms in Africa. Other varieties illustrated by Santschi (1930b) are shown right.

Collingwood (1985), recording it as having been reported from Saudi Arabia, noted that in profile the dorsal outline of the alitrunk is more or less continuous; the gaster with a distinct pattern of 2 or 3 yellow blotches, which appear as a row of 2 or 3 discrete spots on either side; gula hairs numerous; larger species, HW 3.6-3.75 mm.

Baroni Urbani (1972: 124 ff) gave an extensive list of the synonyms and very brief notes on reasons for the synonymies. His primary view appears to have been that the species displays great plasticity in colour patterns across its wide range. Occasional reference is made to variations in size of the major workers, e.g. the head width, but no detailed analysis of dimensions, such as scape length, hind tibia length, size of spicules on the hind tibia (if any) etc., was given.

Listing as in Bolton (1995: 109)

The confusion is well illustrated by Forel (1879a) on {original description}. and {original description}. Emery (1920c) attempted a maculatus-group key; this is at {original description}.

It seems that much of the synonymy stems from Baroni Urbani (1972: 120 ff).

Present subspecies:

African
miserabilis (C. maculatus st. miserabilis n.st., Santschi, 1914d, illustrated, worker) - probably a distinct species {C. maculatus miserabilis}Santschi's (1914d) description is at {original description}. TL 6.5-9.0 mm Guinea
ugandensis (Camponotus (Dinomyrmex) liengmei For. v.ugandensis n.var., Santschi, 1923e: 292, worker) Santschi's (1923e) description is at {original description}. TL 8-8.5 mm; Uganda
Non-African
foveolatus (Camponotus (Myrmoturba) maculatus r. foveolatus, Stitz, 1925: 125, soldier & worker) Stitz's (1925) description is at {original description}. Philippines
Stitz, H. 1925. Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsber. Ges. Naturforsch. Freunde Berlin 1923, 110-136.
humilior (Camponotus maculatus var. humilior, Forel, 1902h: 497, soldier & worker, also subspecies status; Taylor & Brown, 1985: 116) Forel's (1902h) description is at {original description}. Given species status by McArthur & Leys (2006)http://www.oegef.at/MN8_99-110.pdf Australia
Forel, A. 1902. Fourmis nouvelles d'Australie. Rev. Suisse Zool., 10, 405-548
obfuscatus (Camponotus (Myrmoturba) sunudus Em. var. n. obfuscata, Viehmeyer, 1916a: 154, soldier & worker; Bolton, 1995: 114, subspecies of maculatus: current status, no authority given) Viehmeyer's (1916a) description is at {original description}. Singapore
Viehmeyer, H. 1916. Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Arch. Naturgesch. 81(A8), 108-168.
strangulatus (Camponotus maculatus stirps strangulatus, Santschi, 1911e: 129, illustrated, soldier, worker & queen); junior synonym maculatoides, Emery, 1920c: 6, worker) from Madagascar - seems most likely to be related to Camponotus (T.) hova than to maculatus {C. maculatus strangulatus minor}Santschi's (1911e) description is at {original description} Madagascar
subnudus (Camponotus rubripes Drury, razza sunudus n. st., Emery, 1889b: 511, soldier & worker; Forel, 1913k: 125, queen; status changed several times, most recent being as subspecies of maculatus, Stitz, 1925: 124) Emery's (1889b) description is at {original description} Burma
Emery, C. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XX. Formiche di Birmania e del Tenasserim raccolte da Leonardo Fea (1885-87). Ann. Mus. Civ. Stor. Nat. Genova, (2)7(27), 485-520.
sylvaticomaculatus (C.rubripes r maculatus var sylvaticomaculatus, Forel 1886f: 150, worker; Dalla Torre, 1893: 241 - listed name only)
of a very dubious status		 Forel's (1886f) mention of the name (from his 1879 paper (see above) and from Madagascar!) is at {original description}. Madagascar [Bolton (1995) has Greece
- apparently citing Dalla Torre, C. G. de 1893. Catalogus hymenopterorum hucusque descriptorum systematicus et synonymicus. Volumen VII: Formicidae (Heterogyna). Sumptibus Guilelmi Engelmann, Lipsiae. 289 pp. (17 Mb)]


Junior synonyms:

atramentarius (Camponotus maculatus F. var. atramentarius, Forel, 1904b: 379, worker; Forel, 1910f: 26, queen) Forel's (1904b) description is at {original description}. Arnold (1922: 625) gave a translation, this is at {original description}.

"Identical to the type form, but almost entirely black; except the funiculi, base of the tarsi and a narrow band on the gaster segments, plus sometimes the sides of alitrunk partially dark yellow brown. A little more shiny than the type, recalls melanoticus and substitutus. Transition to the type is by all the varieties of colour. Tibiae not setose"; from Ethiopia, collector Dimitriev".

 Ethiopia & South Africa
ballioni (Camponotus maculatus F. r. ballioni n.st., Forel, 1904d: 176, worker & queen) {C. maculatus ballioni frons}Forel's (1904d) description is at {original description}. Arnold (1922: 632) gave a note, this is at {original description}. "Only a darker variety of the true cognatus" South Africa
boera (Camponotus maculatus F. ssp cognatus var boera n. var., Forel, 1910f: 27; name listed ballioni var boera by Santschi, 1925h: 166, soldier, worker & queen) Forel's (1910f) description is at {original description}. Arnold (1922: 632) gave a translation, this is at {original description}. Entirely rusty yellow-red or brownish-yellow colour; shape, sculpture and pilosity agree entirely with cognatus South Africa
cavallus (Camponotus maculatus st. ballioni n.st., Santschi, 1911g: 211, soldier, worker & queen) {Camponotus maculatus cavallus minor}Santschi's (1911g) description of cavallus is at {original description}. TL 9.5-10.5 mm; blackish brown but with extensive reddish-yellow areas, notably the basal two-thirds of the gaster (or of the first gaster segment?), second segment almost completely black; mandibles with some striation; frontal carinae more S-shaped and posteriorly further apart than type maculatus Angola
cluisoides (Camponotus maculatus F. var cluisoides n. var., Forel, 1913h: 354, soldier & worker)

Photomontage derived from images at www.discoverlife.org - originals by Gary Alpert, Harvard University; Kenya specimens.
These specimens are noticeably smaller than the type and are more contrasting black and yellow in the major, with a pallid minor (the scales on the Alpert major seem erroneous).

Also confusing is the labelling, his website now (2008) has this as C. maculatus ugandensis, without a collection label

 {Camponotus maculatus ? cluisoides}Forel's (1913h) description is at {original description}. TL 11.5-15.5 mm; major HL 4.4 HW 4.3; major head strongly enlarged and impressed behind (more than with cluis); base colour black, tibiae and metatarsis dark brown, funiculi and tarsi reddish brown, other light areas yellow, gaster with long lateral patches (as in the type). Kenya
{Camponotus maculatus ? cluisoides minor}
cognata (Formica cognata, Smith, F., 1858b: 35, soldier & worker; Mayr, 1862: 655, queen & male) Smith's (1858b) description is at {original description}. "Extremely like F. maculata, which it may be; the antennae and head as in that species; the legs entirely ferruginous; the abdomen entirely black". South Africa
conakryensis (Camponotus (Myrmoturba) maculatus F. var. conakryensis n., Emery, 1920c: 13, soldier & worker) Emery's (1920c) description is at {original description}. Major TL 14 mm; body black, a little less opaque than var. lohieri; first segment of gaster totally yellow belwo, above with an irregularly triangular black area; pronotum and mesonotum brown; tibia brown Guinea
erythraea (Camponotus (Myrmoturba) maculatus F. var. erythraea n., Emery, 1920c: 14, soldier) Emery's (1920c) description is at {original description}. Santschi (1935b) had {original description} from Kenya. TL 15 mm; colouration as conakryensis but with almost whole of alitrunk dorsum black; head wholly opaque with positions of ocelli distinct.Minor with much narrower head than type maculatus; colouration as in C. aegyptiacus Eritrea
flavifemur (Camponotus (Myrmoturba) maculatus F. var. conakryensis n.v., Santschi, 1937g: 84, worker & queen) Santschi's (1937g) description is at {original description}. Worker similar to lohieri and zempti Sant. but with the femora and tibiae pale yellow contrasting with the red ochre or ferruginous mesopleurosternum and metasternum (as in the other varieties); tibia with brown black aguillons passing into ferruinous in the distal third. Tanzania
flavominor (Camponotus (Myrmoturba) maculatus F. st melanocnemis Sants. v flavominor n.v., Santschi, 1920i: 4, soldier & worker; name in Emery, 1925b: 87, soldier & worker) Santschi's (1920i) description is at {original description}. Size and colour as lohieri but differs from melanocnemis, lohieri and thomensis by the yellow colour of the head and appendages (brown in the others); major propodeum profile almost rectilinear (weakly convex in lohieri) Benin
hannae (Camponotus (Myrmoturba) maculatus F. st hannae n.st., Santschi, 1919a: 349, illustrated, soldier & worker) {C. maculatus hannae minor}Santschi's (1919a) description is at {original description}. Arnold (1926: 294) gave a translation, this is at {original description}. TL 8-12 mm; major coloured like race cavallus but this has a shorter more convex alitrunk; tibiae compressed and without setae Zimbabwe
hieroglyphicus (Camponotus (Myrmoturba) maculatus F. st hieroglyphicus n.st., Santschi, 1917b: 290, soldier, worker & queen) Santschi's (1917b) description is at {original description}. TL major 11.5 mm, HL (with mandibles) 3.5 HW HW 2.5; base colour black but some dark areas, including the mandibles dark brown red; funiculus and part of tibiae reddish, other light areas yellowish; light patches on gaster sharply defined occupying about the borders of the first three segments; gaster shiny, rest of head and body matt, densely reticulo-punctate Angola
intonsus (C. maculatus intonsus n. subsp., Emery 1905d: 29, footnote, soldier & worker) Emery's (1905d) description is on {original description}. Arnold (1922: 632) gave a translation, this is at {original description}. Major TL 12.5 mm, HW 3/1, SL 2.8; gaster of major wholly brown or black, either without patches or with dark indistinct patches; tibiae with ventral setae South Africa
lacteipennis (Formica lacteipennis, Smith F., 1858b: 34, all forms) Smith's (1858b) description is at {original description}. Major TL 9.5 mm, minor 8.5; base colour black; funiculi and legs rufo-testaceous, tarsi darker; head very large deeply impressed behind South Africa
liengmei (Camponotus sexpunctatus n. spec. var. liengmei, Forel, 1894b: 67, soldier & worker; Forel, 1907g: 88, male); given various statuses, this was placed as a subspecies/race of maculatus by Forel (1901h: 69), it was raised to species by Santschi (1923e: 292), then reduced to a junior synonym of maculatus by Baroni Urbani (1972: 125). The distinctive images of the cotype and a fresh specimen from Kenya, persuade me that it merits full species status. {Camponotus maculatus liengmei}Forel's (1894b) description is at {original description}. Arnold (1922: 636) gave a translation, this is at {original description}.
As with Camponotus sexpunctatus, this is a large form, major TL 15-17.3 mm; The photomontage is of a cotype of Camponotus maculatus ssp liengmei. The original photographs, together with enlarged images, are from the MCZ, Harvard University, website at - MCZ link		 Mozambique
liocnemis (Camponotus maculatus liocnemis n. subsp., Emery, 1905d, footnote, soldier & worker; Santschi, 1914e: 38, queen & male, named only) {C. maculatus liocnemis gaster}Emery's (1905d) description is at {original description}. Arnold (1922: 625) gave a translation, this is at {original description}. Smaller than the type form, major TL 10.5 mm, with a smaller head; colouration more contrasting yellow and blcck South Africa
lividior (Camponotus maculatus v. lividior n. var., Santschi, 1911e; 128, all forms) Santschi's (1911e) description is at {original description}. Similar to the type form but minors with the base colour dull greyish yellow Comoro Is
lohieri (Camponotus (Myrmoturba) maculatus st melanocnemis var lohieri, Santschi, 1913c: as variety of maculatus, Emery, 1915g: 22, worker) {lohieri gaster} {C. maculatus lohieri fron}Santschi's (1913c) description as a variety of melanocnemis was - "Worker - head more matt especially towards the posterior angles, and the gaster with a yellow ochre patch on the side of the second segment plus three patches at the base of the first segment, otherwise the gaster is entirely black, as with melanocnemis", from Ivory Coast, Jacqueville, coll R P J H Lohier. Ivory Coast
manzer (Camponotus maculatus subsp. manzer n. subsp., Forel, 1910e: 452, soldier) Forel's (1910e) description is at {original description}. Arnold (1922: 625) gave a translation, this is at {original description}. Major TL 13 mm, head near trapeziform, as lohieri, with sides slightly less convex than the type maculatus; scapes longer, supassing occiput by about two-fifths of their own length; alitrunk less convex, petiole scale strongly biconvex; lateral light patches on first and second gaster segments confluent and none of more posterior segments. Zimbabwe
mathildae (Camponotus maculatus subsp. mathildae n. subsp., Forel, 1910c: 266, soldier, worker & queen) Forel's (1910c) description is at {original description}. Santschi (1914b) had more at {original description}. Arnold (1922) gave a translation which is at {original description}. Tl major 10.5 mm; blck, alitrunk dark brown, legs and mandibles brighter brown; apical segments of gaster yellowish South Africa
melanocnemis (H. Pobeguin, in Santschi, 1911c: 368, soldier; Camponotus (Myrmoturba) maculatus F. stirpsmelanocnemis , Santschi, Santschi, 1915c: 278, worker: raised to subspecies by Forel, 1915, with its junior synonym schultzei, Forel, 1912j: 179, soldier, worker & male) {Camp. maculatus melanocnemis} Santschi (1911c) had the major as - differing from the type (maculatus) by the darker colour of the gaster patches, which are brownish-black; sometimes the gaster is entirely black, the tibiae brown-black; the sculpture is slightly shinier on the posterior angles of the head; the dorsum of the propodeum is slightly longer.
Santschi (1915c) described melanocnemis - this race is very close to the type in general form. The head is a little larger and the frontal carinae slightly further apart, The thorax profile is as arcuate, but somewhat more robust and the propodeum is larger. The posterior tibiae are longer, more than 4 mm. The sculpture is as matt as the type on the head and thorax, but the sides of those parts are generally more shiny. The brown-red pilosity is translucent on the head, otherwise as the type. The overall colour is always darker; with the posterior edges of the thorax and gaster, except for the patches, black or nearly so, although the narrow yellow border persists as in the type; yellow ochre parts of legs are less extended, the tibiae and tarsi are more or less brown black. The patches on the gaster are much more reduced, sometimes absent, The male is black leading to the conclusion that this is a sub-species - in the type the male is yellow with brown patches.
The minor has the head black or dark brown-red; the gaster is black, sometimes basally spotted; the tibiae and dorsal thorax are darker but sometimes ochre in the smallest workers. Head slightly longer than the type and slightly wider anteriorly, with sides straight forward of the eyes and less convex behind, with a distinct occipital border. Thorax profile less convex than type and than in desantii. Petiole scale lower. Specimens from Zaïre - Oubangi, by Rev. P. Augustin; also from Congo, Grand Lahou, by H. Pobéguin. Santschi (1915c) also notes on lohieri (above) as a variety with the patches rather clearer and more apparent, but smaller; except for the base and upper part of the first gastral segment with is more or less yellow ochre, the sculpture is less shiny. Specimens from Ivory Coast, Jacqueville by Lohier; types from Benin, Upper Dahomey, Kouandé, by Desanti; also from Saõ Thomé, by Gravier; and Senegal, by J.de Gaulle.
Santschi's (1915c) description is at {original description}. Arnold (1922) gave a translation/description which is at {original description}. Forel's (1911j) description of schultzei is at {original description}. 		Zaïre
Uganda (schultzei)
The photomontage of what it appears may well be a minor of melanocnemis is of a specimen from the Central African Republic, 2005; {Camponotus maculatus melanocnemis minor}
nubis (Camponotus (Myrmoturba) maculatus (Fabr.) nubis, subsp. nov., Weber, 1943c: 385, illustrated, soldier & worker)

{Camponotus maculatus nubise}Weber's (1943c) description is at {original description}.
Major TL 9.7 mm, no mention of any light patches. Baroni Urbani (1972: 129) appears not to have sighted the Weber type specimen but syninymized it with maculatus as a simple chromatic variety close to melanocnemis not with any obvious morphological differences. The minor specimen below obviously is fairly distinctive

 Sudan (south only)
.

{short description of image}The photomontage of a specimen (below) from Sudan, Bentoe, South Sudan Province, collected by Awatif Omer, 2006, Sudan 13; exactly matches the alitrunk profile drawn by Weber, and has a very similar characters and colour.

 .
radamoides (Camponotus maculatus i.sp., var. radamoides, Forel, 1891b: 213, soldier & worker) Forel's (1891b) description was brief and noted the major and minor workers were similar to the type form but the gaster was entirely black; the occipital angles on the major were reddish; generally smaller - the major with less convex sides to the head and the petiole scale of the minor lower and more obtuse (Andramgoloaka Forest, collector Sikora. Madagascar
sarmentus (Camponotus (Myrmoturba) maculatus F., subsp. sarmentus n., Emery, 1920c: 14, illustrated, soldier & worker) {Camponotus maculatus sarmentus} Emery's (1920c) description is at {original description}. Major TL 13 mm; black, mandibles ferruginous; funiculi, coxae and base of femur reddish-fulvous Kenya
schereri (Camponotus maculatus F., subsp. Schereri n. subsp., Forel, 1911e: 289, worker & queen) Forel's (1911e) description of schereri is at {original description}. Major TL 12-13.5 mm. Hindleg black; gaster with well marked small yellow flecks; head wider than type. Santschi (1915c) also described schereri as - differs from melanocnemis by having more matt sculpture on the sides of the thorax and head. The thorax profile is less convex; and petiole scale less convex in front. Erect hairs a little more abundant on the body, but colour the same. Sometimes the angles of the head are slightly rust coloured, also seen in lohieri. The posterior tibiae are ca. 4 mm, with well developed denticles. Specimens from Liberia, the types of Scherer, from Nebena; southern Nigeria, Olokomeji, by Silvestri; and, Guinea, Kakoulima, by Silvestri. Liberia
semispicatus (C. (Myrmoturba) maculatus st. mathildae var.semispicta, Santschi, 1914b: 130, worker; Emery, 1920c: 5; Menozzi, 1930b: 116, queen) Santschi's (1914b) description is at {original description}. Differs from mathildae by the colour of the sides of the alitrunk and legs which are yellow ochre, that is darker and more like melanconemis Kenya
sudanicus (Camponotus (Myrmoturba) maculatus (Fabr.., sudanicus subsp. nov., Weber, 1943c: 385, soldier & worker; synonymy by Baroni Urbani 1972: 125) {Camponotus maculatus sudanicus} Collated illustration (right) is of a cotype of Camponotus maculatus ssp sudanicus (Weber, 1943c: 385, soldier & worker; synonymy by Baroni Urbani 1972: 125) from Southern Sudan. The original photographs, together with enlarged images, are from the MCZ, Harvard University, website at - MCZ link. Weber's (1943c) description of sudanicus is at {original description}. Major TL 9.1 mm (AL 3.3 mm) body dark brown almost black; smaller than melanocnemis; darker than aegyptiacus and without spots; nest arboreal. Sudan
thomensis (Camponotus (Myrmoturba) maculatus Fab. v.. thomensis n.var., Santschi, 1920i: 3, soldier, worker & male) Santschi's (1920i) description is at {original description}. Major TL 9.5 mm, characterized by the smallness of the lateral patches on the gaster São Tomé I.
tuckeri (Camponotus (Myrmoturba) maculatus F. var. tuckeri nov., Santschi, 1932a: 391, soldier, worker & male) Santschi's (1932a) description is at {original description}. Major TL 13 mm; colour as in the type but with larger patches on the gaster, confluent as a V on the first tergite; base colour black, except fuiniculi, small tarsi, edges of pronotum, mesosternum, metasternum and genae rust. Underside of head with a few hairs. Head longer as with Camponotus aegyptiacus but narrower anteriorly Namibia
zumpti (Camponotus (Myrmoturba) maculatus F. v.. zumpti n.v., Santschi, 1937b: 103, worker) Santschi's (1937b) description of zumpti is at {original description}. Major TL 14 mm; HL 3.8-3.9 HW 3.6-3.8; scape slightly surpasses the occiput; overall colour and light patches as with lohieri, also more shiny Cameroun

Unavailable names:

calceatus (Santschi, 1930b: 76, soldier & worker) {Camponotus (Tanaemyrmex) macualts calceatus}Santschi's (1930b) description is at {original description} Angola
cluis (Forel, 1909b: 67, worker) Forel's (1909b) description is at {original description}. Angola
cataractae (Santschi, 1919b: 239, soldier & worker) Santschi's (1919b) description is at {original description}
Zimbabwe
citinus (Santschi, 1930b; see Bolton, 1995) {C. maculatus citinus}Santschi's (1930b) description is at {original description}
Santschi (1935) noted that in citrinus, probably the correct spelling of what elsewhere is given as citinus, the spots on the gaster are small as in hieroglyphicus Santschi from Benguela, but the latter race is smaller, with the head less impressed posteriorly		 Zaïre
cognatomaculatus (Camponotus rubripes r cognatus var cognatomaculatus, Forel, 1886f: 151, worker; given as Forel, 1889: 255, unavailable name in Bolton 1995: 93 Forel's (1886f) mention of the name (two specimens before him) is at {original description}. Kakoma - Equatorial Africa (there are several possibilities; the name Kakoma appears twice in Congo DR, possibly the most likely candidate
contaminatus (Santschi, 1917b: 291, soldier & worker) Santschi's (1917b) description is at {original description} Zimbabwe
diffusus (Santschi, 1917b: 292, soldier & worker) Santschi's (1917b) description is at {original description} Zimbabwe
georgei (Santschi, 1923e: 290, illustrated, soldier & worker) {Camponotus maculatus georgei} Santschi's (1923e) description is at {original description} Zimbabwe
hansingi (Forel, 1910e: 452, soldier) Forel's (1910e) description is at {original description} Mozambique (also South Africa)
importunoides (Forel, 1914d: 249, soldier & worker) Forel's (1914d) description is at {original description} South Africa
incommoda (Forel, 1914d: 250, soldier & worker) Forel's (1914d) description is at {original description} South Africa
madecassa (Emery, 1905d: 30, footnote, worker) Emery's (1905d) description is at {original description}. Madagascar
pessimus (Camponotus maculatus subspecies miserabilis Santschi variety pessimus, new variety, Wheeler, 1922: 235, soldier & worker)

Wheeler (1922) had - major TL only 6-6.5 mm, minor TL 5-5.5 mm, as miserabilis but much smaller; "the head of the major is distinctly narrower anteriorly, the cheeks being less convex and the frontal carinae are less approximated. Sculpture, pilosity and color very much as in miserabilis" - it seems possible that this is synonymous with Camponotus (Tanaemyrmex) agonius but that is inadequately described; image collated from http://mcz-28168.oeb.harvard.edu/mcz/FMPro?-DB=Image.fm&-Lay=web&-Format=images.htm&Species_ID=21488&-Find

 Zaïre
{short description of image}
Contents Camponotus maculatus main entry
© 2004, 2005, 2006, 2008 - Brian Taylor CBiol FIBiol FRES
11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K.

href="camp_maculatus_variation.htm"